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Euderopus sp (Biphyllidae) [image by K.Darrow]
Euderopus sp.:(Biphyllidae) (image by K.Darrow)

Floyd Shockley, Acting Collections Manager

  • Phone:   202-633-0982
  • Fax:   202-786-2894
  • E-mail Address:
  • Mailing Address:
    Department of Entomology
    Smithsonian Institution
    PO Box 37012, MRC 165
    Washington, DC 20013-7012
  • Shipping Address:
    Smithsonian Institution
    Nat'l Museum of Natural History
    10th & Constitution NW
    Washington, DC 20560-0165
  • Education:
    B.A. Westminster College
    M.S. University of Missouri
    Ph.D. University of Georgia
Floyd Shockley

Duties and Interests:


I am the primary contact for the superfamily Cucujoidea (Coleoptera) (excluding 2 families: Phalacridae and Coccinellidae), handling all specimen loan requests, performing curatorial duties, and hosting visitors. I am also the primary contact for curation, transaction management and hosting scientific visitors for all deactivated groups (~18% of the collection).

Collection Management

As Acting CM, I work on all aspects of collection management, logistics, purchasing and property management for the Department of Entomology, providing complete oversight of the National Insect Collection (100,000+ type specimens, 35 million+ specimens) and interacting with staff from the three agencies that comprise the Combined Entomology department (SI, USDA and DoD).

I am the chair of the Entomology Collections Committee and the Entomology representative on the NMNH Collections, IPM and Safety committees, as well as the 4th floor (East Court) representative on the Entomology Space committee.  I serve as COTR for CM projects, develop new policies and procedures involving the handling and management of specimens, coordinate purchasing and inventory control of collection supplies and equipment, and serve as the primary liaison between the scientific community and Entomology staff.  I am responsible for specimen preparation, sorting, and management of the department specimen backlog, and overseeing/training contractors, interns and volunteers. I perform all pest identification and remediation recommendations for the department IPM program, as well as providing identification support services to the NMNH community and the public, as needed.  I am the primary point of contact for all OFMR, OBM, OCIO and OPS staff operations.   

In addition to conducting many tours of the Entomology collections ranging in audiences from high level dignitaries to small children’s groups, I am also involved in Entomology public educational outreach and social media endeavors, frequently interacting with Education & Outreach and Public Relations staff. I am also responsible for development of new educational materials for tours of the insect collection and special exhibits. 

Systematics Research

My research interests focus on the taxonomy and systematics of the beetle family Endomychidae s.l. (Coleoptera: Cucujoidea), a heterogeneous group of obligate fungus feeders. I reconstruct phylogenetic relationships of Endomychidae using molecular evidence, and use the resulting evolutionary hypotheses to explore patterns of change in physical, behavioral, and ecological characteristics. My expertise further extends to include the alpha-level taxonomy and natural history of the group (especially Neotropical members), and I am particularly interested in comparative morphology of adults and immature stages, as well as fossils.


Shockley, F.W. and Thomas, M. 2015. Notes on the taxonomic identity of Trogosita pusillima Mannerheim, 1843, with transfer from Laemophloeidae and synonymy underHoloparamecus depressus Curtis, 1833 (Endomychidae: Merophysiinae). Pan Pacific Entomologist, 91(3): 278-280

Robertson, J.A., A. Slipinski, M. Moulton, F.W. Shockley, A. Giorgi, N.P. Lord, D.D. McKenna, W. Tomaszewska, J. Forrester, K.B. Miller, M.F. Whiting, and J.V. McHugh.  2015. Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia). Systematic Entomology, 91(3): 278-280.  

Arriaga-Varela, E., W. Tomaszewska, and F.W. Shockley. 2015. Two new species of Epipocus (Coleoptera: Endomychidae) from Mexico and Costa Rica, with a discussion on species groups in Epipocus. The Canadian Entomologist 147 (2): 148-157.

Shockley, Floyd W. and Alekseev, Vitaly I. 2014Glesirhanis bercioi, a new genus and species from Baltic amber (Coleoptera: Endomychidae: Leiestinae) with a checklist and nomenclatural notes regarding fossil Endomychidae.  Zootaxa, 3755(4): 391-400.doi:10.11646/zootaxa.3755.4.5

Szczepanowska, H., F.W. Shockley, D.G. Furth, P. Gentili, D. Bell, P.T. DePriest, M. Mecklenburg, and C. Hawks. 2013.  Effectiveness of entomological collection storage cabinets in maintaining stable relative humidity in a historic museum building.  Collection Forum 27(1): 43-53.

Shockley, F.W., and D. Greenwalt. 2013.  Ptenidium kishenehnicum (Coleoptera: Ptiliidae), a new fossil described from the Kishenehn oil shales, with a checklist of previously known fossil ptiliids.  Proceedings of the Entomological Society of Washington 115(2): 173-181.
Shockley, F.W.  2013.  Advances in Imaging Capabilities in Entomology.  NMNH IT Times 48: 2.

Cline, A.R., and F.W. Shockley.  2012.  A new species of Fallia Sharp (Cucujoidea: Discolomatidae) from the West Indies, with a world checklist for FalliaThe Coleopterists Bulletin 66(2):  93–99.

Shockley, F.W.  2012.  Diversity, Phenology and Natural History of Insects and other Arthropods in The Far Side® by Gary Larson.  American Entomologist 58(2):  74–77.

Shockley, F.W.  2012. First records of Micropsephodes lundgreni Leschen and Carlton (Coleoptera: Endomychidae) in Alabama and Texas, U.S.A.  The Coleopterists Bulletin 66(1): 76–78.

Shockley, F.W.  2011.  Book Review:  Ciegler, J.C.  2010.  The Weevils of South Carolina (Coleoptera: Nemonychidae, Attelabidae, Brentidae, Ithyceridae, and Curculionidae).  Biota of South Carolina.  Volume 6.  Clemson University, Clemson, SC.  Annals of the Entomological Society of America 104(5):  1027-1028.

Stanger-Hall, K.F., F.W. Shockley and R.E. Wilson. 2011. Teaching students how to study: A workshop on information processing and self-testing helps students learn.  CBE-Life Sciences Education 10: 187-198.

pdf Shockley, F.W. and N.J. Vandenberg. 2011. Notes on the taxonomic identity of Bystus hirtulus (Kirsch) and transfer from Endomychidae to Coccinellidae (Coleoptera: Cucujoidea), with designation of a lectotype for Alexia hirtula Kirsch. Zootaxa 2868: 62-68.

Arriaga-Varela, E. and F.W. Shockley. 2010. New records of the Neotropical handsome fungus beetle genus Ephebus Gerstaecker (Endomychidae: Epipocinae) from Mexico and Costa Rica. Dugesiana 17(2): 209-211.

Cline, A.R. and F.W. Shockley.  2010.  10.6. Biphyllidae LeConte, 1861.  Pp. 306-311. In:  Leschen, R.A.B., R.G. Beutel, and J.F. Lawrence (eds.), Handbuch der  Zoologie/Handbook of Zoology. Band/Volume IV. Arthropoda: Insecta Teilband/Part 38. Coleoptera, Beetles. Volume 2. Morphology and Systematics (Polyphaga partim).  W. DeGruyter, Berlin. 786 p.

Shockley, F.W. 2010. Micropsephodes bahamaensis, a new species of Anamorphinae (Coleoptera: Cucujoidea: Endomychidae) from the Bahamas, with a key to the New World genera of Anamorphinae. Insecta Mundi 0112: 1-11.

Shockley, F.W. 2009. Endomychidae. Pp. 109-114. In: Gerlach, J. (ed.), The Coleoptera of the Seychelles Islands.  Pensoft Publishers, Sofia-Moscow.  266 p.

pdf Shockley, F.W. and N.J. Vandenberg. 2009. Catalogue of the primary types of Cerylonidae, Endomychidae and Latridiidae (Coleoptera: Cucujoidea) deposited in the National Museum of Natural History, with additional notes and clarification of the status of several types.  Zootaxa 2229: 1-64.

Shockley, F.W.  2009.  The Systematics Resources Web Portal—The best web resource you’re not using (and may not have even known existed)…  ESA Newsletter 32 (1): 5.

Shockley, F.W. and M.D. Ulyshen.  2009. An interesting deformity in Alobates pennsylvanica (DeGeer), 1775 (Coleoptera: Tenebrionidae).  Journal of Entomological Science 44: 284-286.

Shockley, F.W., K.W. Tomaszewska and J.V. McHugh.  2009.  Review of the natural history of the Handsome Fungus Beetles (Coleoptera: Cucujoidea: Endomychidae).  Insecta Mundi 0072: 1-24.

Shockley, F.W. 2009.  Systematics of Endomychidae (Coleoptera: Cucujoidea): Phylogenetic and Taxonomic Research on a Mycophagous Beetle Family.  Ph.D. Dissertation.  Department of Entomology, University of Georgia. 499 p.

pdf Shockley, F.W., K.W. Tomaszewska and J.V. McHugh.  2009.  An annotated checklist of the handsome fungus beetles of the world (Coleoptera: Cucujoidea: Endomychidae). Zootaxa 1999: 1-113.

Shockley, F.W., M.D. Ulyshen and N.P. Lord.  2008. New distributional records and natural history notes for Micropsephodes lundgreni Leschen and Carlton (Coleoptera: Endomychidae). Coleopterists Bulletin 62(3): 350-352.

Shockley, F.W. and K.W. Tomaszewska.  2007.  First larval description for Symbiotes gibberosus (Lucas) (Coleoptera: Endomychidae).  Annales Zoologici 57(4): 751-755.

Shockley, F.W.  2007.  Stenotarsus nigrivestis, a New Species of Endomychidae (Coleoptera: Cucujoidea) from the Dominican Republic.  Coleopterists Bulletin 61(3): 479–486.

Shockley, F.W.  2006.  Discolomopsis dominicana, a new genus and species of Handsome Fungus Beetle (Coleoptera: Endomychidae) from Dominican Amber.   Insecta Mundi 20(3-4): 1-4.

Shockley, F.W. and K. Murray.  2006.  Discovery of a parasitoid and a predator of batflies (Diptera: Streblidae) at La Selva, Costa Rica.  Biotropica 38(6): 789-790. 

Shockley, F.W. and A.R. Cline.  2004.  A Contribution to the Inventory of Coleoptera of Missouri:  Records from Benton County.  Journal of the Kansas Entomological Society 77 (3): 280-284.

McGraw, R.L., F.W. Shockley, J.F. Thompson, and C.A. Roberts.  2004.  Evaluation of Fifteen Native Legume Species for Forage Yield, Quality, and Seed Production.  Native Plants Journal 5(2):152-159.

Shockley, F.W., R.L. McGraw, and H.E. Garrett.  2004.  Growth and nutrient concentration of two native forage legumes inoculated with Rhizobium and Mycorrhiza in Missouri, USA.  Agroforestry Systems 60:137-142.

McGraw, R.L., F.W. Shockley, and T. Elam.  2003.  Effects of temperature on germination of ten native legume species.  Native Plants Journal 4 (1):5-9.

Cline, A.R., F.W. Shockley, and B. Puttler.  2002.  Description of Surface Swimming by Hypera eximia (Coleoptera: Curculionidae):  Behavioral, Morphological, and Phylogenetic Implications.  Annals of the Entomological Society of America 95 (5): 637-645.

Cline, A.R., F.W. Shockley, and B. Puttler.  2002.  Description of Surface Swimming by Hypera eximia (Coleoptera: Curculionidae):  Behavioral, Morphological, and Phylogenetic Implications.  Annals of the Entomological Society of America 95 (5): 637-645.

McGraw, R.L., D.A. Sleper, and F.W. Shockley.  2002.  Temperature effects on seed development of turf types of tall fescue.  International Herbage Seed Group Newsletter 35:5-8.

Shockley, F.W.  2002.  Instability of dominance hierarchies among male Madagascar hissing cockroaches, Gromphadorhina portentosa (Dictyoptera:  Blaberidae).  Journal of Entomological Science 37 (3):281-282.

Shockley, F.W., E.A. Backus, M. Ellersieck, D. Johnson, and M. McCaslin.  2002.  Glandular-haired alfalfa resistance to potato leafhopper (Homoptera:  Cicadellidae) and hopperburn:  Development of resistance indices.  Journal of Economic Entomology 95 (2):437-447.

Shockley, F.W. and E.A. Backus.  2002.  Repellency to the potato leafhopper (Homoptera:  Cicadellidae) by erect glandular trichomes on alfalfa.  Environmental Entomology 31(1):22-29.

Shockley, F.W.  2000.  Variation in Resistance of Glandular-haired Alfalfa to the Potato Leafhopper (Homoptera: Cicadellidae) and Hopperburn. M.S. Thesis. Department of Entomology, University of Missouri.  60 p.

Electronic Publications:

Shockley, F.W. and A.R. Cline. 2011.  Biphyllidae. False skin beetles, Biphyllid beetles.
In: The Tree of Life Web Project: 

Shockley, F.W. and A.R. Cline. 2009. Discolomatidae. Discolomid beetles, Discolomatid beetles, Mexican hat beetles. 
In: The Tree of Life Web Project:

Shockley, F.W., C.S. Hartley and N. Lord. 2009. Latridiidae. Minute brown scavenger beetles.
In: The Tree of Life Web Project:

Shockley, F.W. 2009. The Handsome Fungus Beetles (Coleoptera: Endomychidae) of Papua Indonesia. In: The Papua Insects Foundation, Taxonomic and faunistic overviews on the insect species living in Papua and West-Papua (Indonesian New Guinea) Project: orders/Coleoptera/Endomychidae/ Endomychidae.htm

Shockley, F.W. 2008. Endomychidae. Handsome fungus beetles.
In: The Tree of Life Web Project:

Shockley, F.W. 2008. Alexiidae Imhoff 1856. Sphaerosoma Samouelle 1819.
In: The Tree of Life Web Project:

Shockley, F.W. 2007. Checklist of the Alexiidae of the World (Coleoptera:  Cucujoidea).

Shockley, F.W. 2007.  A Bibliography of Endomychidae (Coleoptera: Cucujoidea).

Shockley, F.W.  2004.  Strohecker Endomychidae Collection at the Florida State Collection of Arthropods.

Shockley, F.W. and J.V. McHugh.  2003.  The Handsome Fungus Beetles (Coleoptera:  Endomychidae) of Great Smoky Mountains National Park. Endomychids.htm

McHugh, J.V. and F.W. Shockley.  2003.  The Pleasing Fungus Beetles (Coleoptera:  Erotylidae) of Great Smoky Mountains National Park. Erotylids.htm

Blog posts:

SHOCKLEY, F.W. 2012.  Shoo Fly, Don’t Bother Bee!!! Posted: January 4, 2012.

SHOCKLEY, F.W. 2011. Cry Havoc and Let Slip the Bugs of War. Posted:  October 20, 2011.

SHOCKLEY, F.W. 2011. It Takes Two to Tango… or Does It? The Curious Courtship of Collembola (Springtails). Posted: October 11, 2011.

SHOCKLEY, F.W. 2011. Where Have All the Bees Gone?  Posted: October 4, 2011. 

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