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SI Entomology Staff
Warren Steiner, Research Collaborator, Coleoptera
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Until his retirement in January 2010, Warren was responsible for the care and management of the Coleoptera collection and provided research assistance to Terry Erwin.
Special interests include the biosystematics of the Tenebrionidae, Phalacridae, and aquatic beetles; insect collection and identification; biodiversity surveys; zoogeography and dispersal; plant-insect interactions; spread of adventive species; habitat conservation; scientific illustration.
On his retirement in January 2010:
“It will not really be retirement! I have many papers and projects already started, some decades old that will finally be addressed over the coming seasons. They include reviews of New World genera of Tenebrionidae, reports of adventive species, and descriptions of larvae and pupae discovered and reared during many travels and backyard collecting. Much of this information will be incorporated into the co-authored (with Jim Dunford, Dan Young and Mike Thomas) draft of “The Darkling Beetles of the Eastern United States,” a future book and related web site to serve as an identification manual, with images, data on species distributions and life histories.
The Phalacridae will also get renewed attention. My Master’s thesis on this group (review of two small genera for North America) was never published, as was the 1981 review of a larger genus while I was Faculty Research Assistant with U. MD. I plan to collaborate with Matt Gimmel, current student (LSU) with a global view of this family, to get these studies in print. As I have always had one foot in the desert and the other in the creek, water beetles will not be forgotten either. Regional surveys, collecting and preparation of specimens will continue for aquatic insects as well as numerous terrestrial taxa.
The survey of Bahamian Tenebrionidae (and other insects) will progress as long as time and money will allow and permits are granted. Other Caribbean islands beckon also, and I hope to contribute to studies by Mike Ivie and his students on this rich fauna. Genera of flightless darkling beetles are of particular interest and have led me into some amusing predictive biogeography, where in many cases I have suspected the occurrence and discovery of certain new species on selected islands where, when known habitats were searched upon arrival, eureka! Many islands and island groups have yet to be sampled seriously by any entomologist and I hope to get to as many as possible.”
Publications:
Aalbu, R. L., K. Kanda, & W. E. Steiner, Jr. (2009). Opatroides punctulatus Brulle´ now established in California (Coleoptera: Tenebrionidae). The Pan-Pacific Entomologist 85(2):38-42. 
Steiner, W. E., Jr. (2009). The Helopini (Coleoptera: Tenebrionidae) of Virginia. In: S. M. Roble and J. C. Mitchell (eds.) A Lifetime of Contributions to Myriapodology and the Natural History of Virginia: A Festschrift in Honor of Richard L. Hoffman’s 80th Birthday. Virginia Museum of Natural History Special Publication No. 16, Martinsville, VA. Pp. 331-339.
Droege, S., C.A. Davis, W.E. Steiner, Jr. and J. Mawdsley. 2009. The lost micro-deserts of the Patuxent River: using landscape history, insect and plant specimens, and field work to detect and define a unique community. Proceedings of the Entomological Society of Washington 111(1): 132-144.
Henry, T.J., Wheeler, A.G. and Steiner, W.E. (2008) First North American records of Amphiareus obscuriceps (Poppius) (Hemiptera: Heteroptera: Anthocoridae), with a discussion of dead-leaf microhabitats. Proceedings of the Entomological Society of Washington, 110: 402-416.
Steiner, W.E. (2008) Ground beetles (Coleoptera: Carabidae) of Navassa Island, West Indies. Annals of Carnegie Museum, 77, 129-134.
Steiner, W. E., Jr. (2008) A checklist of the darkling
beetles (Insecta: Coleoptera: Tenebrionidae) of
Maryland, with notes on the species recorded from
Plummers Island through the 20th Century. Bulletin of the Biological
Society of Washington 15: 133-140.
Steiner, W.E., Erwin, T.L. and Ward, R.D. (2007) Phloeoxena signata (Dejean): Northern range extensions to Maryland and Tennessee, U.S.A., and the first record for Costa Rica (Coleoptera: Carabidae). The Coleopterists Bulletin, 61, 224-226.
Beutel, R., Balke, M. and Steiner, W.E. (2006) The systematic position of Meruidae (Coleoptera: Adephaga) and the phylogeny of the smaller aquatic adephagan beetle families. Cladistics, 22, 102-131.
Nearns, E.H. and Steiner, W.E. (2006) A new species of Plectromerus Haldeman (Coleoptera: Cerambycidae: Curiini), from Navassa Island, Greater Antilles. Zootaxa, 1163, 61-68.
Steiner, W.E.,Jr. (2006) New species of darkling betles (Coleooptera: Tenebrionidae) from San Salvador Island, Bahamas. Zootaxa, 1158, 1-38.
Hoffman, R. L. and Steiner, W.E.,Jr. (2005) Jadera haematoloma, another insect on its way north (Hemiptera: Rhopalidae). Banisteria 26: 7-10.
Spangler, P.J. and Steiner, W.E. (2005) A new aquatic beetle family, Meruidae, from Venezuela (Coleoptera: Adephaga). Systematic Entomology, 29, 339-357. [28M]
Steiner, W.E.,Jr. (2005) Two new species of three-horned Hypogena from Peru (Coleoptera: Tenebrionidae). Annales Zoologica, 55, 571-574.
Steiner, W.E.,Jr. (2005) Studies on the darkling beetles (Coleoptera: Tenebrionidae) known from Grand Bahama Island, with descriptions of new species of Branchus and Adlina. Proceedings of the Entomological Society of Washington, 107, 441-459.
Staines, C., W. E. Steiner, Jr. and J. McCann (2005) Surprise in Seth State Forest. National Wetlands Newsletter 27, 7-10.
Steiner, W. (2004) New distribution records for Eleates depressus (Randall) in the southeastern United States (Coleoptera: Tenebrionidae; Bolitophaginae) and notes on these occurrences. Banisteria, 22, 52-53.
Steiner, W. (2004) New distribution records and recent spread of Hymenorus farri Campbell (Coleoptera: Tenebrionidae; Alleculinae) to Florida and in the Caribbean Region. Proceedings of the Entomological Society of Washington, 106, 739-746.
Steiner, W. (2004) The first records of Bycrea villosa Pascoe (Coleoptera: Tenebriondae) in the United States, Central America and Colombia with notes on its associations with leaf-cutting ants. The Coleopterists Bulletin, 58, 329-334.
Steiner, W.E. (2004) The genus Trachyscelis (Coleoptera: Tenebrionidae) in North America: synonymy, origin, distribution, and decline. The Coleopterists Bulletin, 58, 335-343.
Steiner, W.E. and Bouchard, P. (2004) First Descriptions of Coelometopini Pupae (Coleoptera: Tenebrionidae) from Australia, Southeast Asia and the Pacific Region with Comments on Phylogenetic Relationships and Anti-Predator Adaptations. Systematic Entomology, 29, 101-114.
Triplehorn, C.A., Steiner, W.E. and Wappes, J.E. (2004) Notes on the Genus Centronopus Solier (Coleoptera: Tenebrionidae). The Coleopterists Bulletin, 58, 245-246.
Spangler, P.J. and Steiner, W.E., Jr. (2003) A new species of water scavenger beetle Coelostoma (s. str.) tina (Coleoptera: Hydrophilidae: Sphaeridiinae), from Kenya, eastern Africa. Proceedings of the Entomological Society of Washington, 105, 1-8.
Steiner, W.E. (2003) An Immigrant Darkling Beetle New to Hawaii (Coleoptera: Tenebrionidae). Bishop Museum Occasional Papers.Records of the Hawaii Biological Survey for 2001-2002, Part 2: Notes., 40-42.
Steiner, W.E., Hoffman, R.L. and Roble, S.M. (2003) Thirteen Additions to the Known Beetle Fauna of Virginia (Coleoptera: Scirtidae, Bothrideridae, Cleridae, Tenebrionidae, Melyridae, Callirhipidae, Cerambycidae, Chrysomelidae). Banisteria, 20, 53-61.
Steiner, W.E., Staines, C.L., Mccann, J.M. and Hellman, J.L. (2003) The Seth Forest Water Scavenger Beetle, a New Species of Hydrochus (Coleoptera: Hydrophiloidea: Hydrochidae) from the Chesapeake-Delmarva Region. The Coleopterists Bulletin, 57, 101-114.
Alarie, Y., Spangler, P.J. and Steiner, W.E., Jr. (2002) Larval morphology of Agabetes Crotch (Coleoptera: Adephaga: Dytiscidae) : The hypothesis of sister-group relationship with the subfamily Laccophilinae revisted. The Coleopterists Bulletin, 56, 547-567.
Steiner, W.E. (2002) Phalacridae Leach 1815. In: American Beetles, vol 2. pp. 335-337.
Steiner, W.E. (2000) What Flightless Beetles in Eastern U.S. "microdeserts" Tell us about Climate History and Species Distribution After the Ice Age. Abstracts, 27th Annual Natural Areas Conference.Managing the Mosaic: Connecting People and Natural Diversity.Natural Areas Association (2000) 25.
Steiner, W.E. (2000) An Entomological Survey of Navassa Island, with Notes on Species Richness and Endemism. Abstracts, 27th Annual Natural Areas Conference.Managing the Mosaic: Connecting People and Natural Diversity.Natural Areas Association (2000), 39.
Steiner, W.E. (2000) Records and Habitats of the "Rare Click Beetle," Cerophytum pulsator (Haldeman), in Virginia and Maryland (Coleoptera: Cerophytidae). Banisteria, 15, 43-45.
Steiner, W.E., Gifford, M.E. and Powell, R. (2000) Relationship of Diet and Prey Availability in Aristellger cochranae, a Gecko from Navassa Island, West Indies. Caribbean Journal of Science, 36, 323-326.
Steiner, W. E. (1999) Flightless beetles in Appalachian “deserts”: studies on the distribution and localized habitats of some species of Tenebrionidae (Coleoptera). Virginia Museum of Natural History Special Publication. 7:125-144.
Steiner, W.E. and Flint, Oliver S., Jr. (1999) Dragonflies of Navassa Island. Argia, 11, 19-21.
Steiner, W.E., Phillips, K.T. and Triplehorn, C.A. (1998) A New Species of Central American Platydema (Coleoptera: Tenebrionidae) Established in Florida. The Coleopterists Bulletin, 52, 291-296.
Steiner, W.E. (1997) Aquatic Beetles (Coleoptera) of Moapa Warm Springs, Nevada. Bulletin of the North American Benthological Society, 14, 177.
Steiner, W.E. (1996) Hydrobiomorpha casta (Say) in Virginia (Coleoptera: Hydrophilidae). Banisteria, 7, 53-55.
Steiner, W.E. (1996) Microcrypticus (Coleoptera: Tenebrionidae) in Hawaii, with Notes on Distribution and Probable Origin. Records of the Hawaii Biological Survey for 1995, Part 2: Notes. Bishop Occasional Papers, 46, 35-37.
Steiner, W.E. (1995) Structures, Behavior, and Diversity of the Pupae of Tenebrionidae (Coleoptera). Pp. 503-539 In: Pakaluk, James; Slipinski, Stanislaw Adam [Eds]. Biology, phylogeny, and classification of Coleoptera: papers celebrating the 80th birthday of Roy A. Crowson. Volume 1. Muzeum i Instytut Zoologii PAN, Warsaw, Poland. 1995: i-xii, 1-558.
Steiner, W.E. (1992) "Ironclad Beetles," the Family Zopheridae, in Maryland: Notes on the Natural History and Distribution of Phellopsis obcordata (Kirby). The Maryland Naturalist, 35, 25-30.
Steiner, W.E. (1991) Branchus whiteheadi, New Species, from Texas, with Notes on the Genus Branchus (Coleoptera: Tenebrionidae). Proceedings of the Entomological Society of Washington, 93, 425-432.
Steiner, W.E. (1988) Larval Stages and Biology of Eastern U.S. Epitragini (Coleoptera: Tenebrionidae), with the Discovery of a New Form of Pupal "Gin-Trap." [Abstract]. Proceedings fo the XVIII International Congress of Entomology, Vancouver, Canada, 51.
Steiner, W.E. and Singh, B.P. (1987) Redescription of the Ergot Beetle, Acylomus pugetanus Casey, with Immature Stages and Biology (Coleoptera: Phalacridae). Proceedings of the Entomological Society of Washington, 89, 744-758.
Mathis, W.N. and Steiner, W.E. (1985) An Adventive Species of Bracydeutera Loew in North America (Diptera: Ephydridae). Journal of the New York Entomological Society, 94, 56-61.
Steiner, W.E. (1984) A Review of the Biology of Phalacrid Beetles. , 424-445.
Steiner, W.E., Krysan, J.L., Branson, T.F. and Schroeder, F.W. (1984) Elevation of Diabrotica sicuanica (Coleoptera: Chrysomelidae) to the Species Level with Notes on the Altitudinal Distribution of Diabrotica Species in the Cuzco Department of Peru. Entomological News, 95, 91-98.
Froeschner, R.C. and Steiner, W.E. (1983) Second Record of South American Burrowing Bug, Scaptocoris castaneus Perty (Hemiptera: Cydnidae) in the United States. Entomological News, 94, 176.
Spangler, P.J. and Steiner, W.E. (1983) New Species of Water Beetles of the Genera Elmoparnus and Pheneps from Suriname (Coleoptera: Dryopidae; Psephenidae). Proceedings of the Entomological Society of Washington, 85, 826-839.
Steiner, W.E. (1983) Notes on Centronopus opacus LeConte from Oklahoma and Arkansas (Coleoptera: Tenebrionidae). The Coleopterists Bulletin, 37, 90-91.
Steiner, W.E. (1982) Poecilocrypticus formicophilus Gebien, a South American Beetle Established in the United States (Coleoptera: Tenebrionidae). Proceedings of the Entomological Society of Washington, 84, 232-239.
Steiner, W.E. (1981) Brathinus nitidus in Maryland, with Notes on Biology (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 35, 233-234.
Steiner, W.E. (1981) A New Species of Trox from Peru, with Notes on the Biology and Distribution of Trox aricensis Gutierrez (Coleoptera: Scarabaeidae). The Coleopterists Bulletin, 35, 205-210.
Steiner, W.E. (1981) A New Genus and a New Species of Laccophiline Water Beetle from Ecuador (Coleoptera: Dytiscidae). The Pan-Pacific Entomologist, 57, 251-259.
Steiner, W.E. and Anderson, J.J. (1981) Notes on the History of Spanglerogyrus albiventris Folkerts, with a New Distribution Record (Coleoptera: Gyrinidae). The Pan-Pacific Entomologist, 57, 124-132.
Steiner, W.E. and Perkins, P.D. (1981) Two New Peruvian Species the Riffle Beetle Genus Xenelmis (Coleoptera: Elmidae). The Pan-Pacific Entomologist, 57, 306-312.
Steiner, W.E. (1980) Laccobius minutoides (Coleoptera: Hydrophilidae) in Alabama. The Coleopterists Bulletin, 34, 392.
Steiner, W.E. (1980) Additional Distribution Records for Onthophagus taurus (Coleoptera: Scarabaeidae). The Coleopterists Bulletin, 34, 332.
Steiner, W.E. (1980) Notes on the Habitats and Distribution of Rhyssemus scaber (Coleoptera: Scarabaeidae). The Coleopterists Bulletin, 34, 377-378.
Steiner, W.E. (1980) A New Tribe, Genus, and Species of Cossyphodine from Peru (Coleoptera: Tenebrionidae). Proceedings of the Entomological Society of Washington, 82, 384-395.
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