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Smithsonian National Museum of Natural History
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Trixoscelis desserta (photos by K. Darrow)
_Trixoscelis sp. (Diptera)

Wayne N. Mathis, Curator Emeritus of Diptera

  • Phone:   202-633-1014
  • Fax:   202-786-2894
  • E-mail Address:   mathisw@si.edu
  • USPS Address:
    Smithsonian Institution
    PO Box 37012, MRC 169, Rm. CE-619
    Washington, DC 20013-7012
  • Public Carrier Address:
    Smithsonian Institution
    National Museum of Natural History
    10th & Constitution NW
    Washington, DC 20560-0169
  • Education:
    BS Brigham Young
    PhD Oregon State
Wayne Mathis

Related Websites:
http://www.nadsdiptera.org/index.htm
http://www.inhs.uiuc.edu/cee/FLYTREE/flyothercollab.htm


Research Interests:

For over 40 years, I have focused on the systematics and evolution of higher Diptera (true flies), especially families in the Ephydroidea and Opomyzoidea. My research primarily employs the methodology of cladistic analysis to unravel, reconstruct, and document phylogenetic relationships and how these relationships impinge on historical biogeography. The phylogenetic investigations and analyses run the gamut from species-level revisions to basal lineages of superfamilies. Documentation includes comprehensive systematics studies at all levels of the classification, faunistic studies, catalogs and conspectuses. Although my comprehensive studies are global, I have focused field work in the neotropics, especially the Caribbean, and on islands, such as Sri Lanka, the Seychelles, New Zealand, and the barrier islands of Belize.

During my tenure at the Smithsonian Institution, I have had the pleasure of working with numerous postdoctoral, graduate, and undergraduate fellows and interns, including Dr. Amnon Freidberg (1979 1980), Dr. Kumar Ghorpadé (1983 1984), Dr. Martha Condon (1987 1989), Dr. Jon K. Gelhaus (1989-1990), Dr. Gregory W. Courtney (1990-1992), Dr. Marion Kotrba (1995-1996), Dr. Darlene D. Judd (1996-1998), Dr. Volker Hollmann (1997-1998), Dr. Stephen D. Gaimari (1999-2000), Dr. Masahiro Sueyoshi (2004-2005), Dr. James F. Edmiston (1989, 2009), Dr. Richard G. Robbins (1990), Dr. Ilka C. Feller (1989-1994), Dr. Suely Marques (1992), Dr. Alessandra Rung (1996-2004), Dr. Peter H. Kerr (1997-2005), Dr. Brian Wiegmann (1989), Dr. Thomas Pape (1987-1988), Dr. Navin Rao (1994), Dr. Kevin Holston (1995), Dr. Luciane Marinoni (1991, 2002), Dr. Isaac S. Winkler (2002), Ms. Rachael Eden (1996), Dr. Torsten Dikow (2000), Dr. Michelle D. Trautwein (2001), Mr. Jin Zuyin (1987-1988). My collaboration with Dr. Tadeusz Zatwarnicki (University of Opole, Poland) over the last 25 years and with George A. Foster over the last 15 years has been especially productive.

My collection responsibilities at the Smithsonian include over 750,000 specimens (pinned, alcohol, and slides) in over 20 families of Diptera.

Publications:

Mathis, W., and Zatwarnicki, T. 2013. A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae). Zookeys, 363: 1-161. doi:10.3897/zookeys.363.6482

Foster, George A. and Mathis, Wayne N. 2012. A revision of the nearctic species of the genus Trixoscelis Rondani (Diptera: Heleomyzidae: Trixoscelidinae). Smithsonian Contributions to Zoology, 637: 1-128.

Huang, Yiau-Min, Mathis, Wayne N. and Wilkerson, Richard C. 2012. Subgeneric Position of Aedes dufouri Hamon with Notes on the Subgenus Levua Stone and Bohart (Diptera: Culicidae). Proceedings of the Entomological Society of Washington, 114(2): 194-204. doi:10.4289/0013-8797.114.2.194

Mathis, Wayne N. and Freidberg, Amnon. 2012. Periscelis stuckenbergi sp. n., the first record of the genus from the Afrotropical Region (Diptera: Periscelididae: Periscelidinae). African Invertebrates, 53(1): 231-238.

Mathis, Wayne N. and Marinoni, Luciane. 2012. A conspectus on the Canacidae (Diptera) of Brazil. ZooKeys, 162: 59-92. doi:10.3897/zookeys.162.2370

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2012. Review of the shore-fly genus Oedenopiforma (Diptera: Ephydridae). The Canadian Entomologist, 144(1): 81-92. doi:10.4039/tce.2012.9

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2012. Revision of New World Species of the Shore-fly Subgenus Allotrichoma Becker of the Genus Allotrichoma with Description of the Subgenus Neotrichoma (Diptera, Ephydridae, Hecamedini). ZooKeys, 161: 1-101. doi:10.3897/zookeys.161.2016

Mathis, Wayne N., Steinly, Bruce A., Jr., Buckingham, Gary R. and Deonier, Bryan C. 2012. Dick L. Deonier 1936-2011 Obituary. Proceedings of the Entomological Society of Washington, 114(1): 169-172.

Mathis, Wayne, Rung, Alessandra and Kotrba, Marion. 2012. A revision of the genus Planinasus Cresson (Diptera, Periscelididae). ZooKeys, 225: 1-83. doi:10.3897/zookeys.225.3721

Zhang, Junhua, Yang, Ding and Mathis, Wayne N. 2012. A review of the species of Rhynchopsilopa Hendel from China (Diptera, Ephydridae). Zookeys, 216: 23-42. doi:10.3897/zookeys.216.3224

Mathis, Wayne N. and Marinoni, Luciane. 2011. A review of Parydra (Diptera: Ephydridae) from Brazil. Zoologia, 28(4): 505-512. doi:10.1590/S1984-46702011000400012

Mathis, W.N. and Sueyoshi, M. 2011. New species of the genus Cyamops Melander from New Zealand (Diptera, Periscelididae, Stenomicrinae). ZooKeys114: 28-40.

Rung, A. and Mathis, W.N. 2011. A Revision of the Genus Aulacigaster Macquart (Diptera: Aulacigastridae). Smithsonian Contributions to Zoology, 633: 1-132.

Huang, Y.-M., Mathis, W.N. and Wilkerson, R.C. 2010. Coetzeemyia, a new subgenus of Aedes, and a redescription of the holotype female of Aedes (Coetzeemyia) fryeri (Theobald) (Diptera: Culicidae). Zootaxa, 2638: 1-24.

Mathis, Wayne N. 2010. Canacidae (surf flies). In: Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, Norman E. and Zumbado, M. A., Manual of Central American Diptera. Ottawa: NRC Research Press, pp.1115-1119.

Mathis, Wayne N. 2010. Coelopidae (kelp flies). In: Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, Norman E. and Zumbado, M. A., Manual of Central American Diptera. Ottawa: NRC Research Press, pp.1013-1016.

Mathis, Wayne N. 2010. Diastatidae (diastatid flies). In: Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, Norman E. and Zumbado, M. A., Manual of Central American Diptera. Ottawa: NRC Research Press, pp.1207-1210.

Mathis, Wayne N. 2010. Ephydridae (shore flies). In: Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, Norman E. and Zumbado, M. A., Manual of Central American Diptera. Ottawa: NRC Research Press, pp.1211-1233.

Mathis, Wayne N. 2010. Nannodastiidae (nannodastiid beach flies). In: Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, Norman E. and Zumbado, M. A., Manual of Central American Diptera. Ottawa: NRC Research Press, pp.1235-1239.

Mathis, Wayne N. 2010. Tethinidae (beach flies). In: Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, Norman E. and Zumbado, M. A., Manual of Central American Diptera. Ottawa: NRC Research Press, pp.1109-1114.

Mathis, Wayne N. and Rung, Alessandra. 2010. Periscelididae (periscelid flies). In: Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, Norman E. and Zumbado, M. A., Manual of Central American Diptera. Ottawa: NRC Research Press, pp.1087-1092.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2010. New Species and Taxonomic Clarifications for Shore Flies from the Delmarva States (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 112(1): 97-128. doi:10.4289/0013-8797.112.1.97

Munari, Lorenzo and Mathis, Wayne N. 2010. World Catalog of the Family Canacidae (including Tethinidae) (Diptera), with keys to the supraspecific taxa. Zootaxa, 2471: 1-84.

Myers, Janet and Mathis, Wayne N. 2010. Hollis (Holly) Barton Williams 1951-2009. Proceedings of the Entomological Society of Washington, 112(3): 481-482.

Zatwarnicki, Tadeusz and Mathis, Wayne N. 2010. A Revision of the Nivea Group of the Shore-fly Genus Ditrichophora Cresson (Diptera: Ephydridae). Transactions of the American Entomological Society, 136(3-4): 199-215.

Kotrba, Marion and Mathis, Wayne N. 2009. The Internal Female Reproductive Tract of the Enigmatic Genus Risa (Diptera: Schizophora: Ephydroidea) and its Phylogenetic Implications. Proceedings of the Entomological Society of Washington, 111(3): 627-640.

Mathis, Wayne N. and Mathis, Dianne. 2009. Species of the Shore-Fly Genus Glenanthe Haliday (Diptera: Ephydridae) from New Mexico. Proceedings of the Entomological Society of Washington, 111(2): 454-463.

Mathis, Wayne N., Knutson, Lloyd V. and Murphy, William L. 2009. A New Species of Snail-Killing Fly of the Genus Dictya Meigen from the Delmarva States (Diptera: Sciomyzidae). Proceedings of the Entomological Society of Washington, 111(4): 785-794.

Mathis, Wayne N., Zatwarnicki, Tadeusz and Kubatova-Hirsova, Hana. 2009. A revision of the shore-fly genus Philotelma Becker (Diptera: Ephydridae). Insect Systematics & Evolution, 40(2): 121-158. doi:10.1163/187631209X416714

Zhang, J., Yang, D. and Mathis, W.N. 2009. A New Species of the Shore-Fly Genus Oedenopiforma Cogan from the Oriental Region, with an Updated Key to the Species (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 111(1): 199-203.

Foster, G. A. and Mathis, Wayne N. 2008. Review of the genus Tethina Haliday (Diptera : Canacidae : Tethininae) from western North America. Proceedings of the Entomological Society of Washington, 110(2): 300-330.

Foster, G. A. and Mathis, Wayne N. 2008. A review of the Tethininae (Diptera: Canacidae) from the Galapagos Islands. Proceedings of the Entomological Society of Washington, 110(3): 743-752.

Mathis, Wayne N. 2008. Two new Neotropical genera of the shore-fly tribe Ephydrini Zetterstedt (Diptera: Ephydridae). Zootaxa, 1874: 1-15.

Mathis, Wayne N. and Mathis, Dianne. 2008. Shore flies (Diptera: Ephydridae) of Plummers Island. In: Brown, J. W., The Invertebrate fauna of Plummers Island, Maryland. Contribution to the Natural History of Plummers Island, Maryland.. Washington, D. C.: Bulletin of the Biological Society of Washington, pp.173-191.

Mathis, Wayne N. and Ohishi, H. 2008. Revision of the shore-fly genus Eremomusca Mathis (Diptera : Ephydridae) with description of a new species from Japan. Proceedings of the Entomological Society of Washington, 110(1): 67-78.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2008. A Revision of the Nearctic Species of Hecamedoides Hendel (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 110(4): 1012-1027.

Brake, Irina and Mathis, Wayne N. 2007. Revision of the genus Australimyza Harrison (Diptera: Australimyzidae). Systematic Entomology, 32(2): 252-275.

Edmiston, J. F. and Mathis, Wayne N. 2007. New Zealand species of the shore-fly genus Nostima Coquillett (Diptera : Ephydridae). Zootaxa, 1661: 1-16.

Mathis, Wayne N. and Foster, G. A. 2007. Canacidae (Diptera) from the Delmarva States. Proceedings of the Biological Society of Washington, 120(4): 387-428.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2007. A revision of the species of the genus Dichaeta Meigen (Diptera: Ephydridae). Annales Zoologici, 57(4): 783-822.

Zatwarnicki, Tadeusz and Mathis, Wayne N. 2007. A revision of the Palearctic species of the shore-fly genus Discomyza Meigen (Diptera : Ephydridae). Insect Systematics & Evolution, 38(3): 241-266.

Zhang, J. J., Yang, D. and Mathis, Wayne N. 2007. Species of the genus Zeros Cresson (Diptera : Ephydridae) from China. Proceedings of the Entomological Society of Washington, 109(4): 872-879.

Marinoni, Luciane and Mathis, Wayne N. 2006. A cladistic analysis of the Neotropoical genus Sepedonea Steyskal (Diptera: Sciomyzidae). Zootaxa, 1236: 37-52.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2006. A review of the New World species of the shore-fly genus Leptopsilopa Cresson (Diptera: Ephydridae). Annales Zoologica, 56(1): 85-138.

Mathis, Wayne N., Castro, B. A., Wasy, M. O. and Zatwarnicki, Tadeusz. 2006. Discovery of the South American rice miner (Diptera: Ephydridae) in the Southeastern United States. Annals of the Entomological Society of America, 99(6): 999-1005.

Edmiston, J. F. and Mathis, Wayne N. 2005. A revision of the New World species of the shore-fly subgenus Nostima Coquillett (Genus Philygria Stenhammar; Diptera: Ephydridae). Smithsonian Contributions to Zoology, 623: 1-108. doi:10.5479/si.00810282.623

Edmiston, J. F. and Mathis, Wayne N. 2005. A review of two Nearctic species of the shore-fly genus Philygria Stenhammar, P. debilis Loew and P. nigrescens (Cresson) (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 107(1): 7-20.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2005. Revision of New World species of shore fly genus Discomyza Meigen (Diptera: Ephydridae. Annals of the Entomological Society of America, 98(4): 431-443.

Mathis, Wayne N. and Zumbado, M. A. 2005. Description of Scatella savegre, a new species from Costa Rica in the triseta group (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 107(2): 386-391.

Rung, A., Mathis, Wayne N. and Papp, L. 2005. Curiosimusca, gen. nov., and three new species in the family Aulacigastridae from the Oriental Region (Diptera: Opomyzoidea). Zootaxa, 1009: 21-36.

Zhang, J., Yang, D. and Mathis, Wayne N. 2005. A new genus and species of Ephydridae (Diptera) from the Oriental Region. Zootaxa, 1040: 31-43.

Mathis, Wayne N. and Rung, Alessandra. 2004. Redescription of Diopsosoma primum Malloch (Diptera: Periscelididae). Revista Brasileira de Entomologia, 48(3): 303-309.

Mathis, Wayne N. and Tadeusz, Zatwarnicki. 2004. Tronamyia, a new genus of shore-flies and a phylogenetic reassessment of the tribe Lipochaetini Becker (Diptera: Ephydridae). Annals of the Entomological Society of America, 97(2): 259-270.

Mathis, Wayne N. and Tadeusz, Zatwarnicki. 2004. A review of the species of Hyadina Haliday (Diptera: Ephydridae) occurring in Israel. Israel Journal of Entomology, 34: 35-57.

Mathis, Wayne N. and Tadeusz, Zatwarnicki. 2004. A revision of the shore-fly genus Cressonomyia Arnaud (Diptera: Ephydridae), with comments on species that have been excluded. Proceedings of the Entomological Society of Washington, 106(2): 249-279.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2004. Description of a new species of Hyadina Haliday from southern California (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 106(3): 705-716.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2004. A review of two Nearctic shore-fly species in the genus Psilopa Fallén that were included in the genus Cressonomyia Arnaud (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 106(3): 639-648.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2004. A revision of the shore-fly genus Trimerogastra Hendel (Diptera: Ephydridae). Pages 89-108. In N. L. Evenhuis and K. Y. Kaneshiro, eds., D. Elmo Hardy Memorial Volume. Contributions to the Systematics and Evolution of Diptera. Bishop Museum Bulletins in Entomology, 12: 89-108.

Mathis, Wayne N., Zatwarnicki, T. and John, W. M. M. 2004. A review of unreported shore-fly genera of the tribe Scatellini from the New Zealand subregion (Diptera: Ephydridae). Zootaxa, 622: 1-27.

Steyskal, George C., Thompson, F. Christian, Mathis, Wayne N. and Knutson, Lloyd. 2004. The type species of Ilione (Diptera: Sciomyzidae). Studia Dipterologica, 10(2): 559-564.

Sueyoshi, Masahiro and Mathis, Wayne N. 2004. A new species of Cyamops Melander, 1913 (Diptera: Periscelididae) from Japan and a review of the Japanese Periscelididae. Proceedings of the Entomological Society of Washington, 106(1): 74-84.

Cheng, Lanna and Mathis, Wayne N. 2003. Marine insects of Guam: Heteroptera and Diptera. Micronesica, 35-36: 514-522.

Foster, George A. and Mathis, Wayne N. 2003. A revision of the genera Pelomyia Williston and Masoniella Vockeroth (Diptera: Tethinidae). Smithsonian Contributions to Zoology, 619: 1-63. doi:10.5479/si.00810282.619

Mathis, Wayne N. and Michelle, D. Trautwein. 2003. A revision and phylogenetic study of Lipochaeta Coquillett (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 105(1): 30-49.

Mathis, Wayne N. and Winkler, Isaac S. 2003. A review of the Neotropical species of Brachydeutera Loew (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 105(2): 407-424.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2003. Shore flies of the Republic of Seychelles (Diptera: Ephydridae). Annales Zoologici (Warszawa), 53(4): 585-650.

Dikow, Torsten and Mathis, Wayne N. 2002. A revision and phylogenetic study of Actocetor Becker (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 104(2): 249-290.

Freidberg, Amnon and Mathis, Wayne N. 2002. The Periscelididae (Diptera) of Israel, with description of a new species of Stenomicra Coquillett. Israel Journal of zoology, 48: 43-51.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 2002. A phylogenetic study of the tribe Dryxini Zatwarnicki (Diptera:Ephydridae). Smithsonian Contributions to Zoology, 617: 1-101. doi:10.5479/si.00810282.617

Kirk-Spriggs, A.H., Ismay, J.W., Ackland, M., Rohácek, J., Mathis, W.N., Foster, G.A., Pape, T., Cranston, P S. and Meier, R. 2001. Inter-tidal Diptera of southwestern Africa (Chironomidae, Canacidae, Chloropidae, Milichiidae, Tethinidae, Ephydridae, Sphaeroceridae, Coelopidae, Sarcophagidae and Anthomyiidae). Cimbebasia, 17: 85-135.

Papp, L. and Mathis, W.N. 2001. A review of the family Nannodastiidae (Diptera). Proceedings of the Entomological Society of Washington, 103(2): 337-348.

Zatwarnicki, Tadeusz and Mathis, Wayne N. 2001. A generic classification of the tribe Discocerinini (Diptera: Ephydridae). Annales Zoologici, 51(1): 1-51.

Baptista, Alessandra R. P. and Mathis, Wayne N. 2000. Notes on the genus Cyamops Melander (Diptera: Periscelididae), including description of ten new species. Proceedings of the Entomological Society of Washington, 102(3): 481-506.

Foster, George A. and Mathis, Wayne N. 2000. Notes on the Neotropical species of Tethina Haliday (Diptera: Tethinidae). Proceedings of the Entomological Society of Washington, 102(33): 542-548.

Marinoni, Luciane and Mathis, Wayne N. 2000. A cladistic analysis of Sciomyzidae Fallén (Diptera). Proceedings of the Biological Society of Washington, 113(1): 162-209.

Mathis, Wayne N. and Edmiston, J. F. 2000. A revision of the shore-fly genus Lemnaphila Cresson (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 102: 652-677.

Mathis, Wayne N. and Grimaldi, David A. 2000. The first Beckeriella Williston from the Afrotropical Region: two new species from Madagascar (Diptera: Ephydridae). In: Goodman, S. M., A floral and faunal inventory of the Parc National de Marojejy, Madagascar: with reference to elevational variation.. Chicago: Fieldiana: Zoology 97, pp.105-115.

Mathis, Wayne N. 1999. A Review of the Beach-Fly Genus Isocance Mathis (Diptera: Canacidae). Proceedings of the Entomological Society of Washington, 101(2): 347-358.

Foster, George A. and Mathis, Wayne N. 1998. A Revision of the Family Tethinidae (Diptera) from the Caribbean, Gulf of Mexico, and Bermuda. Proceedings of the Entomological Society of Washington, 100(4): 601-632.

Mathis, Wayne N. and Family Periscelididae. 1998. Family Canacidae. In: Papp, László and Darvas, Béla, Contributions to a Manual of Palaearctic Diptera (with special reference to flies of economic importance). Vol. 3. Higher Brachycera. Budapest: Science Herald, pp.251-257.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 1998. Family Ephydridae. In: Papp, László and Darvas, Béla, Contributions to a Manual of Palaearctic Diptera (with special reference to flies of economic importance). Vol. 3. Higher Brachycera. Budapest: Science Herald, pp.537-570.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 1998. A Review of the West Indian Species of Mimapsilopa Cresson (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 100(1): 7-24.

Mathis, Wayne N., Papp, László. 1998. Family Periscelididae. In: Papp, László and Darvas, Béla, Contributions to a Manual of Palaearctic Diptera (with special reference to flies of economic importance). Vol. 3. Higher Brachycera. Budapest: Science Herald, pp.285-294.

Papp, László and Mathis, Wayne N. 1998. Family Nannodastidae. In: Papp, László and Darvas, Béla, Contributions to a Manual of Palaearctic Diptera (with special reference to flies of economic importance). Vol. 3. Higher Brachycera. Budapest: Science Herald, pp.309-314.

Feller, Ilka C. and Mathis, Wayne N. 1997. Primary Herbivory by Wood-Boring Insects along an Architectural Gradient of Rhizophora mangle. Biotropica, 29(4): 440-451.

Mathis, Wayne N. 1997. A Review of the Hoguei Group of Paracanace Mathis and Wirth (Diptera: Canacidae). Memoirs of the Entomological Society of Washington, 18: 140-148.

Mathis, Wayne N. 1997. A Revision of Neotropical Ditricohphora Cresson (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 99(4): 697-704.

Mathis, Wayne N. 1997. A Review of the Shore-Fly Genus Diphuia Cresson (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 99(1): 28-36.

Mathis, Wayne N. 1997. Shore Flies of the Belizean Cays (Diptera: Ephydridae). Smithsonian Contributions to Zoology, 592: 1-77. doi:10.5479/si.00810282.592

Baptista, A. R. P. and Mathis, Wayne N. 1996. A New Species of Cyamops Melander (Diptera: Periscelididae) from Brazil, with Distributional Notes on Another Species. Proceedings of the Entomological Society of Washington, 98(2): 245-248.

Mathis, Wayne N. 1996. Australian beach flies (Diptera: Canacidae). Proceedings of the Biological Society of Washington, 109(2): 326-348.

Mathis, Wayne N. and Munari, L. 1996. World Catalog of the Family Tethinidae (Diptera). Smithsonian Contributions to Zoology, 584: 1-27. doi:10.5479/si.00810282.584

Mathis, Wayne N. 1995. Studies of Gymnomyzinae (Diptera: Ephydridae), VI: A Revision of the Genus Glenathe Haliday from the New World. Smithsonian Contributions to Zoology, 567: 1-26. doi:10.5479/si.00810282.567

Mathis, Wayne N. 1995. Shore Flies of the Galapagos Islands (Diptera: Ephydridae). Annals of the Entomological Society of America, 88(5): 627-640.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 1995. World Catalog of Shore Flies (Diptera: Ephydridae). Memoirs on Entomology, International, Associated Publishers, 4: 1-423.

Rao, N. and Mathis, Wayne N. 1995. A Revision of the Shore-fly Genus Rhysophora Cresson (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 97(3): 666-688.

Baptista, A. and Mathis, Wayne N. 1994. A Revision of the New World Cyamops Melander (Diptera: Periscelididae). Smithsonian Contributions to Zoology, 563: 1-24. doi:10.5479/si.00810282.563

Mathis, Wayne N. 1994. A Revision of the Shore-Fly Genus Clasiopella Hendel (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 96(3): 454-465.

Mathis, Wayne N. and Freidberg, Amnon. 1994. A Revision of the Neartic Aulacigaster MacQuart with Notes on A. Leucopeze (Meigen) from the Paleartic Region (Diptera: Aulacigastridae). Proceedings of the Entomological Society of Washington, 96(4): 583-598.

Zatwarnicki, Tadeusz and Mathis, Wayne N. 1994. Phylogeny and Classification of the Genus Scatophila Becker (Diptera: Ephydirdae). Annales de la Societe Entomologique de France, 29(4): 351-370.

Evenhuis, Neal L., Mathis, Wayne N. and Thompson, F. Christian. 1993. Comment on the Proposal to Conserve Oecothea Haliday in Curtis, 1836 and to Designate Helomyza fenestralis Fallen, 1820 as the Type Species (Insect, Diptera). Bulletin of Zoological Nomenclature, 50(3): 235-236.

Grimaldi, D. A. and Mathis, Wayne N. 1993. Fossil Periscelididae (Diptera). Proceedings of the Entomological Society of Washington, 95(3): 383-403.

Mathis, Wayne N. 1993. Studies of Gymnomyzinae (Diptera: Ephydridae), IV: A Revision of the Shore-Fly Genus Hecamede Haliday. Smithsonian Contributions to Zoology, 541: 1-46. doi:10.5479/si.00810282.541

Mathis, Wayne N. 1993. A New Species and Subgenus of Periscelis Loew from Australia (Diptera: Periscelididae). Journal of the Australian Entomological Society, 32: 13-19.

Mathis, Wayne N. 1993. A Revision of the Shore-Fly Genera Hostis Cresson and Paratissa Coquillett (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 95(1): 21-47.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 1993. A Revision of the Shore-Fly Genus Chlorichaeta Becker (Diptera: Ephydridae). Tijdschrift voor Entomologie, 136(1): 55-76.

Mathis, Wayne N., Zatwarnicki, Tadeusz and Krivosheina, M. G. 1993. Studies of Gymnomyzinae (Diptera: Ephydridae), V: A Revision of the Shore-Fly Genus Mosillus Latreille. Smithsonian Contributions to Zoology, 548: 1-38. doi:10.5479/si.00810282.548

Mathis, Wayne N. 1992. The First Shore Fly of the Genus Glenanthe Haliday from the Australasian Region (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 94(1): 78-82.

Mathis, Wayne N. 1992. World Catalog of the Beach-Fly Family Canacidae (Diptera). Smithsonian Contributions to Zoology, 536: 1-18. doi:10.5479/si.00810282.536

Mathis, Wayne N. and Papp, L. 1992. A New Genus of Periscelididae (Diptera) from the Neotropics. Proceedings of the Biological Society of Washington, 105(2): 366-372.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 1992. Ephydridae Zetterstedt, 1837 (Insecta, Diptera): Proposed Precedence over Gymnomyzidae Latreille, 1829. Bulletin of Zoological Nomenclature, 49(2): 133-136.

Mathis, Wayne N. 1991. Studies of Gymnomyzinae (Diptera: Ephydridae), II: A Revision of the Shore-Fly Subgenus Pseudohecamede Hendel of the Genus Allotrichoma Becker. Smithsonian Contributions to Zoology, 522: 1-28. doi:10.5479/si.00810282.522

Mathis, Wayne N. 1991. Classification of the Shore Flies (Diptera: Ephydridae) Past, Present, and Future. In: Proceeding of the II International Congress of Dipterology. pp.209-228.

Mathis, Wayne N. and Edmiston, J. F. 1991. S. W. Williston's Species of Ephydridae (Diptera) from St. Vincent (West Indies). Proceedings of the Biological Society of Washington, 104(4): 816-839.

Mathis, Wayne N. and Freidberg, Amnon. 1991. Review of the Afrotropical Beach Flies of the Tribe Canacini and Subfamily Nocticanacinae (Diptera: Canacidae). Proceedings of the Entomological Society of Washington, 93(1): 70-85.

Freidberg, A. and Mathis, W.N. 1990. A New Species of Craspedoxantha and a Revised Phylogeny for the Genus (Diptera: Tephritidae). Proceedings of the Entomological Society of Washington, 92(2): 325-332.

Mathis, Wayne N. 1990. A Revision of the Shore-Fly Genus Diphuia Cresson (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 92(4): 746-756.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 1990. A Revision of the Western Paleartic Species of Athryoglossa (Diptera: Ephydridae). Transactions of the American Entomological Society, 116(1): 103-133.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 1990. Taxonomic Notes on Ephydridae (Diptera). Proceedings of the Biological Society of Washington, 103(4): 891-906.

Mathis, Wayne N. 1989. 103. Family Canacidae. In: Catalog of the Diptera of the Australasian and Oceanian Regions, special publication 86. pp.669-670.

Mathis, Wayne N. 1989. 53. Family Cypselosomatidae. In: Catalog of the Diptera of the Australasian and Oceanian Regions, special publication 86. pp.463

Mathis, Wayne N. 1989. 84. Family Coelopidae. In: Catalog of the Diptera of the Australasian and Oceanian Regions, special publication 86. pp.563-564.

Mathis, Wayne N. 1989. 99. Family Ephydridae. In: Catalog of the Diptera of the Australasian and Oceanian Regions, special publication 86. pp.639-649.

Mathis, Wayne N. 1989. 56. Family Pseudopomyzidae. In: Catalog of the Diptera of the Australasian and Oceanian Regions, special publication 86. pp.470

Mathis, Wayne N. 1989. Diptera (Insecta) or True Flies of the Pitcairn Group (Ducie, Henderson, Oeno, and Pitcairn Islands). Atoll Research Bulletin, 327: 1-15. doi:10.5479/si.00775630.327.1

Mathis, Wayne N. 1989. A Review the Beach Flies of the Carribean and Gulf of Mexico (Diptera: Canacidae). Proceedings of the Biological Society of Washington, 102(3): 590-608.

Mathis, Wayne N. and Jin, Z. Y. 1989. A Review of the Shore Fly Genus Polytrichophora Cresson from Asia (Diptera: Ephydridae). Proceedings of the Biological Society of Washington, 102(2): 434-446.

Mathis, Wayne N. and Sasakawa, M. 1989. 102. Family Tethinidae. In: Catalog of the Diptera of the Australasian and Oceanian Regions, special publication 86. pp.86

Mathis, Wayne N. 1988. The First Record of the Genus Procanace Hendel from North America, with the Description of a New Species (Diptera: Canacidae). Proceedings of the Entomological Society of Washington, 90(3): 329-333.

Mathis, Wayne N. 1988. Beach Flies of the Republic of Seychelles (Diptera: Canacidae). Bulletin of the Washington Biological Society, 8: 22-29.

Mathis, Wayne N. 1988. First Record of the Shore-Fly Genus Placopsidella Kertacsz from North America (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 90(3): 334-337.

Mathis, Wayne N. and Jin, Z. Y. 1988. A Review of the Asian Species of the Genus Lamproscatella Hendel (Diptera: Ephydridae). Proceedings of the Biological Society of Washington, 101(3): 540-548.

Mathis, Wayne N. and Zatwarnicki, Tadeusz. 1988. Studies on the Shore Fly Tribe Dagini (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 90(1): 106-121.

Mathis, Wayne N. and Deeming, J. C. 1987. A Revision of the Shore Fly Genus Elephantinosoma Becker (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 89(3): 562-570.

Wirth, W. W., Mathis, Wayne N. and Vockeroth, J. R. 1987. 98. Ephydridae. In: Manual of the Families and Genera of Neartic Diptera Volume 2. pp.1027-1047.

Freidberg, Amnon and Mathis, Wayne N. 1986. Studies of Terelliinae (Diptera: Tephritidae): A Revision of the Genus Neaspilota Osten Sacken. Smithsonian Contributions to Zoology, 439: 1-75. doi:10.5479/si.00810282.439

Mathis, Wayne N. 1986. Studies of Psilopinae (Diptera: Ephydridae), I: A Revision of the Shore Fly Genus Placopsidella Kertacsz. Smithsonian Contributions to Zoology, 430: 1-30. doi:10.5479/si.00810282.430

Mathis, Wayne N. 1986. A Revision of the Subgenus Eremotrichoma Soika of the Shore Fly Genus Allotrichoma Becker (Diptera: Ephydridae). Isreal Journal of Entomology, 19: 127-139.

Mathis, Wayne N. and Hogue, C. L. 1986. Description of a New Species of the Shore Fly Genus Diedrops (Diptera: Ephydridae) from Colombia. Contributions in Science (Los Angeles County Museum), 377: 21-26.

Mathis, Wayne N. and Steiner, Warren E. 1986. An Adventive Species of Brachydeutera Loew in North America (Diptera: Ephydridae). Journal of the New York Entomological Society, 94(1): 56-61.

Freidberg, Amnon and Mathis, Wayne N. 1985. On the Feeding Habits of Rhynchopsilopa (Diptera: Ephydridae). Entomophaga, 30(1): 13-21.

Mathis, Wayne N. 1985. A New Psilopine Genus and Species from Isreal with a Recharacterization of the Tribe and Key to Other Psilopine Genera of the Middle East (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 87(2): 375-380.

Mathis, Wayne N. 1985. Studies of Parydrinae (Diptera: Ephydridae), II: A Revision of the Shore Fly Genus Pelinoides Cresson. Smithsonian Contributions to Zoology, 410: 1-46. doi:10.5479/si.00810282.410

Mathis, Wayne N. and Ghorpade, K. D. 1985. Studies of Parydrinae (Diptera: Ephydridae), I: A Review of the Genus Brachydeutera Loew from the Oriental, Australian, and Oceanian regions. Smithsonian Contributions to Zoology, 406: 1-25. doi:10.5479/si.00810282.406

Mathis, Wayne N. and Steiner, Warren E. 1985. An Adventive Species of Bracydeutera Loew in North America (Diptera: Ephydridae). Journal of the New York Entomological Society, 94(1): 56-61.

Mathis, Wayne N. 1984. A Revision of the Genus Asmeringa Becker (Diptera: Ephydridae). Isreal Journal of Entomology, 17: 67-79.

Mathis, Wayne N. 1984. A Revision of the Shore Fly Genus Homalometopus Becker (Diptera: Ephydridae). Proceedings of the Biological Society of Washington, 97(2): 253-264.

Mathis, Wayne N. 1984. Notes on the Shore Fly Genus Diedrops (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 86(2): 349-353.

Mathis, Wayne N. 1983. Notes on Brachydeutera Loew (Diptera: Ephydrididae) from North America. Entomological news, 94(3): 177-180.

Mathis, Wayne N. 1983. A revision of the genus Dagus Cresson (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 85(4): 717-726.

Mathis, Wayne N. 1982. Ephydridae. In: Aquatic Biota of Mexico, Central America and the West Indies. pp.469-472.

Mathis, Wayne N. 1982. Canacidae of Isreal, with a Revison of the Paleartic Species of the Genus Canace Haliday (Diptera). Entomologica Scandinavica, 13: 57-66.

Mathis, Wayne N. 1982. Description of a New Species of Nocticanace Malloch (Diptera: Canacidae) from Sri Lanka, with Notes on Two Related Species. Proceedings of the Entomological Society of Washington, 84(3): 421-425.

Mathis, Wayne N. 1982. Studies of Ephydrinae (Diptera: Ephydridae), VI: Review of the Tribe Dagini. Smithsonian Contributions to Zoology, 345: 1-30. doi:10.5479/si.00810282.345

Mathis, Wayne N. 1982. Studies of Canacidae (Diptera), I: Suprageneric Revision of the Family, with Revisions of New tribe Dynomiellini and New Genus Isocanace. Smithsonian Contributions to Zoology, 347: 1-21. doi:10.5479/si.00810282.347

Mathis, Wayne N. 1982. Studies of Ephydrinae (Diptera: Ephydridae), VII: Revision of the Genus Setacera Cresson. Smithsonian Contributions to Zoology, 350: 1-57. doi:10.5479/si.00810282.350

Mathis, Wayne N. and Freidberg, Amnon. 1982. New Beach Flies of the Genus Xanthocanace Hendel with a Review of the Species from the Western Paleartic (Diptera: Canacidae). Memoirs of the Entomological Society of Washington, 10: 97-104.

Mathis, Wayne N. 1981. Ephydridae. In: Aquatic Biota of Tropical South America, Part 1: Arthropoda. pp.312-316.

Mathis, Wayne N. 1981. Proposed Use of the Plenary Powers to Grant Precedence to the Family-Group Name Ephydridae over Hydrelliidae (Diptera). Bulletin of Zoological Nomenclature, 38(3): 201-204.

Mathis, Wayne N. and Simpson, K. W. 1981. Studies of Ephydrinae (Diptera: Ephydridae), V: Systematics, Phylogeny, and Natural History of the Genera Cirrula Cresson and Dimecoenia Cresson in North America. Smithsonian Contributions to Zoology, 329: 1-51. doi:10.5479/si.00810282.329

Mathis, Wayne N. and Wirth, W. W. 1981. Studies of Ephydrinae (Diptera: Ephydridae), IV: Revision of the Australian Species of Subgenus Neoscatella Malloch. Smithsonian Contributions to Zoology, 325: 1-27. doi:10.5479/si.00810282.325

Mathis, Wayne N. and Wirth, W. W. 1981. Hydrellia tritici (Diptera: Ephydridae), New to Hawaii. Proceedings of the Hawaiian Entomological Society, 23(3): 371-373.

Thompson, F. C. and Mathis, Wayne N. 1981. Haliday's Generic Names of Diptera First Publised in Curtis' A Guide to British Insects (1837). Journal of the Washington Academy of Sciences, 70(2): 80-89.

Thompson, F. C. and Mathis, Wayne N. 1981. Review of: A Catalogue of Diptera of the Afrotropical Region. British Museum (Natural History). Bulletin of the Entomological Society of America, 27(3): 230-231.

Mathis, Wayne N. 1980. Studies of Ephydrinae (Diptera: Ephydridae), III: Revisions of Some Neotropical Genera and Species. Smithsonian Contributions to Zoology, 303: 1-50. doi:10.5479/si.00810282.303

Mathis, Wayne N. and Freidberg, Amnon. 1980. A New Species of Nesopsilopa from Florida (Diptera: Ephydridae). Florida Entomologist, 63(3): 279-281.

Mathis, Wayne N. and Steyskal, G. S. 1980. A Revision of the Genus Oedoparena Curran (Diptera: Dryomyzidae: Dryomyzinae). Proceedings of the Entomological Society of Washington, 82(3): 349-359.

Mathis, Wayne N. and Zack, R. S. 1980. A New Species of Limnella Malloch (Diptera: Ephydridae) from Mt. Rainier, Washington. Entomological news, 91(3): 65-67.

Deonier, D. L., Mathis, Wayne N. and Regensburg, J. T. 1979. Natural History and Life-cycle Stages of Notiphila carinata (Diptera: Ephydridae). Proceedings of the Biological Society of Washington, 91(4): 798-814.

Mathis, Wayne N. 1979. Ecuadorian Parascatella (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 81(4): 602-610.

Mathis, Wayne N. 1979. Review of: Record of My Life-Work in Entomology, 1903-1904, by C. R. Osten Sacken, with an appreciation and introductory preface by K. G. V. Smith. Proceedings of the Entomological Society of Washington, 81(2): 330-331.

Mathis, Wayne N. 1979. Studies of Ephydrinae (Diptera: Ephydridae), II: Phylogeny, Classification, and Zoogeography of Neartic Lamproscatella Hendel. Smithsonian Contributions to Zoology, 295: 1-41. doi:10.5479/si.00810282.295

Mathis, Wayne N. 1979. Studies of Notiphilinae (Diptera: Ephydridae), I: Revision of the Neartic Species of Notiphila Fallen, Excluding the Caudata Group. Smithsonian Contributions to Zoology, 287: 1-111. doi:10.5479/si.00810282.287

Mathis, Wayne N. 1979. Ephydrinae (Diptera: Ephydridae): A new perspective. In: in: First Symposium on the Systematics and Ecology of Ephydridae (Diptera). pp.47-60.

Mathis, Wayne N. and Wirth, W. W. 1979. Beach flies of Madagascar (Diptera: Canacidae). Annals of the Natal Museum, 23(3): 785-796.

Wirth, W. W. and Mathis, Wayne N. 1979. A Review of the Ephydridae (Diptera) Living in Thermal Springs. In: in: First Symposium on the Systematics and Ecology of Ephydridae (Diptera). pp.21-45.

Mathis, Wayne N. 1978. New Synonymy in Ephydridae (Diptera). Entomological news, 89-num-2&3): 105-106.

Mathis, Wayne N. 1978. A Revision of the Neartic Species of Limnellia Malloch (Diptera: Ephydridae). Proceedings of the Biological Society of Washington, 91(1): 250-293.

Mathis, Wayne N. 1978. A Synopsis of Neotropical Eleleides Cresson (Diptera: Ephydridae). Proceedings of the Entomological Society of Washington, 80(4): 465-472.

Mathis, Wayne N. and Shewell, G. E. 1978. Studies of Ephydrinae (Diptera: Ephydridae), I: Revisions of Parascatella Cresson and the Triseta group of Scatella Robineau-Desvoidy. Smithsonian Contributions to Zoology, 285: 1-44. doi:10.5479/si.00810282.285

Mathis, Wayne N. and Wirth, W. W. 1978. Paracanace New Genus near Canaceoides Cresson, Three New Species, and Notes on their Classification (Diptera: Canacidae). Proceedings of the Entomological Society of Washington, 80(4): 524-537.

Mathis, Wayne N. 1977. A Revision of the Genus Rhysophora Cresson with a Key to Related Genera (Diptera: Ephydridae). Proceedings of the Biological Society of Washington, 90(4): 921-945.

Mathis, Wayne N. 1977. Key to the Neotropical Genera of Parydrinae with a Revision of the Genus Eleleides Cresson (Diptera: Ephydridae). Proceedings of the Biological Society of Washington, 90(3): 553-565.

Mathis, Wayne N. 1977. A New Synonym and Lectotype Designation in Ephydridae (Diptera). Proceedings of the Entomological Society of Washington, 79(4): 537

Mathis, Wayne N. and Wirth, W. W. 1977. New Synonymy in Ephydridae (Diptera). Proceedings of the Entomological Society of Washington, 79(4): 520

Mathis, Wayne N. and Wirth, W. W. 1977. A New Genus of Psilopine flies (Diptera: Ephydridae) with Notes on its Relationships. Proceedings of the Entomological Society of Washington, 79(1): 63-74.

Mathis, Wayne N. and Wirth, W. W. 1976. A New Neotropical Shore Fly Genus with Two New Species (Diptera: Ephydridae). The Pan-Pacific Entomologist, 52(2): 126-132.

Mathis, Wayne N. 1975. A Systematic Study of Coenia and Paracoenia (Diptera: Ephydridae). Great Basin Naturalist, 35(1): 65-85.

Mathis, Wayne N. 1974. Lectotype Designation for Notiphila varia Jones (Diptera: Ephydridae). Entomological news, 85: 64-65.

Mathis, Wayne N. 1973. A Review of the Genus Borboropsis (Diptera: Heleomyzidae). The Pan-Pacific Entomologist, 49(4): 373-377.

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