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Photo of a carabid beetle, Cicindis horni Bruch
Cicindis horni
Bruch
The Fairy Shrimp Hunting Beetle

Terry Erwin, Curator of Coleoptera
Chairman, Department of Entomology

  • Research Office :   202-633-1022
  • Chair's Office: 202-633-1033
  • Fax:   202-786-3141
  • E-mail Address:   erwint@si.edu
  • Mailing Address:
    Smithsonian Institution
    PO Box 37012, MRC 187, Rm. CE-723
    Washington, DC 20013-7012
  • Shipping Address:
    Smithsonian Institution
    National Museum of Natural History
    10th & Constitution NW
    Washington, DC 20560-0187
  • Education:
    AA Vallejo Junior college
    BS & MS San Jose State
    PhD University of Alberta, Edmonton
  • Post docs:
    MCZ, Harvard
    University at Lund, Sweden

Research Interests:

Terry Lee Erwin was born on December 1, 1940, in California’s wine country at St. Helena. His father was a "tin-knocker" and race car driver in the California circuit and his mother a government clerk. Terry spent his youth trout fishing in the High Sierra with his maternal grandfather. As a teenager, with prodding from his father, he began building hot cars and was a founding member and later President of the California Conquistadores, a hot rod club in the Bay Area. He put himself through college by working the Atomic Submarines at Mare Island Naval Base where he was a helper in the "Asbestos" department. At this time, he made a decision that following in his father’s foot steps was not a good idea and became a serious student under the guidance of the Coleopterist, J. Gordon Edwards, at San Jose State College. With Gordon’s enthusiasm and guidance, Terry solved the intractable taxonomy of the North American Bombardier Beetles and went on to write his dissertation on the world fauna under the mentorship of George E. Ball at the University of Alberta, Canada., finishing in 1969.

Having decided that he wanted to work under the three greatest (living) carabidologists, the first being Prof. Ball, Terry obtained postdoctoral fellowships at the Museum of Comparative Zoology at Harvard with Philip J. Darlington Jr. and at Lund University, Sweden with Carl H. Lindroth. However, a position opened at the (then) United States National Museum in the Department of Entomology upon the retirement of Oscar Cartwright. With the support of Washington Biologists Field Club (WBFC) members Paul Spangler (Coleopterist) and Karl Krombien (Chairman of Entomology), Terry was able to accept the position AND within two months took a year-long sabbatical to Sweden. He returned in 1971, to take up the reins as the second Coleopterist with the Department. While in Sweden, the chairmanship of the Department changed to WBFC member Paul D. Hurd, who saw on his desk a proposal left by Terry with Dr. Krombien to obtain funding for studies of the California carabid beetles. Hurd, having learned of monies for studies in Central America, crossed out the "California" and wrote in "Panama". To Terry’s great surprise, he arrived home from Sweden to find out he was on the next plane to the Canal Zone. That was the beginning of a lifetime career on studies of biodiversity in neotropical forests.

With the publication of a small paper (1 1/2 pages) in 1981 on the beetle fauna of a Panamanian tree species, Terry created a cottage industry in canopy studies and in trying to estimate the number of species on the planet. He had hypothesized in that paper that there were perhaps were as many as 30,000,000 species rather than the number described at the time of 1 million – an order of magnitude difference. A lot of people got excited, especially those in the Conservation business because if true, we were losing a lot more species than previously thought.

Terry continues his studies of biodiversity in the western Amazon Basin, at present in Ecuador, and does taxonomic studies of Carabid beetles.

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FULL PUBLICATION LIST

Erwin, T.L., 1965. Trachykele opulenta Fall (Buprestidae) emerging from furniture. Pan-Pacific Entomologist 41(1):68.

Erwin, T.L., 1965. A revision of Brachinus of North America: Part I. The California species (Coleoptera: Carabidae). The Coleopterists Bulletin 19(1):1-19, 17 figs.

Erwin, T.L., 1966. Cyphon sp. (Helodidae) reared from tree hole. Pan-Pacific Entomologist 42(1):68.

Erwin, T.L., 1966. Brachinus pallidus Erwin, a carabid beetle with an entomophagous larva. Pan-Pacific Entomologist 42(1):73.

Erwin, T.L., 1967. Bombardier beetles (Coleoptera: Carabidae) of North America: Part II. Biology and behavior of Brachinus pallidus Erwin in California. The Coleopterists Bulletin 21(2):41-55, 15 figs.

Erwin, T.L., 1969. A new bombardier beetle in Canada (Coleoptera: Carabidae). Opuscula entomologica 34(3):283-286.

Erwin, T.L., 1969. Preliminary diagnoses of five new North American Brachinus (Coleoptera: Carabidae). Opuscula entomologica 34(3):287-288.

Ball, G. E. & Erwin, T.L., 1969. A synopsis of the Tribe Loricerini (Coleoptera: Carabidae). Canadian Journal of Zoology 47(5):877-907, 62 figs., 4 tables.

Erwin, T.L., 1970. A reclassification of bombardier beetles and a taxonomic revision of the North and Middle American species (Carabidae: Brachinida). Quaestiones entomologicae 6:4-215, 460 figs., 16 tables [Dissertation].

Erwin, T.L., 1970. The nearctic species of the Genus Leistus Frolich (Coleoptera: Carabidae). Pan-Pacific Entomologist 46:111-119, 8 figs.

Erwin, T.L., 1970. Unique structures in members of Tachys sensu lat. (Coleoptera: Carabidae). Pan-Pacific Entomologist 46:231-232, 2 figs.

Erwin, T.L., 1970. A description of the larva of Thyce harfordi Casey (Coleoptera: Scarabaeidae). Psyche 77(1):50-53, 10 figs.

Erwin, T.L., 1971. Notes and corrections to a reclassification of bombardier beetles (Carabidae, Brachinida). Quaestiones entomologicae 7:281.

Erwin, T.L., 1971. A teratomorphic carabid beetle with notes on polymorphic asymmetry of the gular region in the same population (Coleoptera: Carabidae). Pan-Pacific Entomologist 47(4):303.

Erwin, T.L., and L. J. Erwin, 1971. A guide to the ground-beetle collection in the Muséum National d'Histoire Naturelle, Paris (Coleoptera: Carabidae). Multilithed by SI Duplicating Unit for limited distribution. 22 pp.

Erwin, T.L., 1971. Fossil tachyine beetles from Mexican and Baltic Amber with notes on a new synonymy of an extant group (Col., Carabidae). Ent. Scand. 2(3):233-236, 5 figs.

Erwin, T.L., 1972. Two new genera of bembidiine carabid beetles from Australia and South America with notes on their phylogenetic and zoogeographic significance (Coleoptera). Breviora No. 383:1-19.

Erwin, T.L., 1972. The lost type of Brachinus sallei Chaudoir (Coleoptera: Carabidae). Pan-Pacific Entomologist 48(1):63-64.

Erwin, T.L., 1972. Observations on the life history of bombardier beetles: Mechanism of egg-hatching (Coleoptera: Carabidae). Pan-Pacific Entomologist 48(1):64.

Erwin, T.L., 1972. Trechus obtusus Erichson in California (Coleoptera: Carabidae). The Coleopterists Bulletin 26(2):42.

Erwin, T.L., 1972. Om biologien hos bombarderbiller (Col., Carabidae, Brachinida). Ent. Meddr. 40:65-71.

Erwin, T.L., & G.E. Ball, 1972. Classification of the Ovipennis and Trifaria groups of Nebria Latreille (Coleoptera: Carabidae: Nebriini). Proceedings of the Biological Society of Washington, 85 (7):77-108.

Erwin, T.L., 1973. Phylogenetic, zoogeographic, and bio-systematic studies of certain carabid ground beetles. American Philosophical Society Year Book 1972, (1973), pp. 362-363.

Erwin, T.L., 1973. Studies of the subtribe Tachyina (Coleoptera: Carabidae: Bembidiini), Part I: A revision of the Neotropical genus Xystosomus Schaum. Smithsonian Contributions to Zoology No. 140, 39 pp.

Erwin, T.L., 1973. Nomenclatorial notes on the Tachyini (Coleoptera: Carabidae). Proceedings of the Entomological Society of Washington 75(1):125.

Erwin, T.L., 1973. Carabid beetles, mountain tops, and trees. Proceedings of the Entomological Society of Washington 75(1):127.

Erwin, T.L., 1973. A supplement to the bombardier beetles of North and Middle America: New records for Middle America (Coleoptera: Carabidae). The Coleopterists Bulletin 27(2):79-82.

Erwin, T.L., 1974. Studies of the subtribe Tachyina (Coleoptera: Carabidae: Bembidiini), Part II: A revision of the New World-Australian Genus Pericompsus LeConte. Smithsonian Contributions to Zoology No. 162, 96 pp.

Erwin, T.L., 1974. Studies of the subtribe Tachyina (Coleoptera: Carabidae: Bembidiini) Supplement A: Lectotype designations for New World species, two new genera, and notes on generic concepts. Proceedings of the Entomological Society of Washington 76(2):123-155.

Erwin, T.L., 1974. The genus Coptocarpus Chaudoir of the Australian Region with notes on related African species (Coleoptera: Carabidae: Oodini). Smithsonian Contributions to Zoology No. 184, 25 pp.

Erwin, T.L., 1974. The ground beetle components of the panamanian fauna, pp.124-128. In, Rubinoff, R. W., 1973. Environmental Monitoring and Baseline Data. Smithsonian Institution Environmental Science Program.

Erwin, T.L., 1975. Studies of the subtribe Tachyina (Coleoptera: Carabidae: Bembidiini), Part III: Systematics, phylogeny, and zoogeography of the genus Tachyta Kirby. Smithsonian Contributions to Zoology No. 208, 68 pp.

Erwin, T.L., 1975. The ground beetle types of Max Liebke in the Smithsonian Institution, Washington, D. C. (Coleoptera: Carabidae). The Coleopterists Bulletin 29(4):267-268.

Erwin, T.L., 1975. Relationships of predaceous beetles to tropical forest wood decay. Part I. Descriptions of the immature stages of Eurycoleus macularis Chevrolat (Carabidae: Lebiini). The Coleopterists Bulletin 29(4):297-300.

Erwin, T.L., 1976. A case of homonymy in the Tachyina (Coleoptera: Carabidae: Bembidiini). The Coleopterists Bulletin 30(1):94.

Erwin, T.L., and L. J. M. Erwin, 1976. Relationships of predaceous beetles to tropical forest wood decay. Part II. The natural history of Neotropical Eurycoleus macularis Chevrolat (Carabidae: Lebiini) and its implications in the evolution of ectoparasitoidism. Biotropica 8(4):215-224.

Erwin, T.L., 1977. Studies of the subtribe Tachyina (Coleoptera: Carabidae: Bembidiini), Supplement B: Lectotype designation of Masoreus ridiculus Schaufuss and transfer of this species to Polyderis. The Coleopterists Bulletin 31(1):90.

Erwin, T.L., D.R. Whitehead & G.E. Ball, 1977. Family 4. CARABIDAE, The Ground Beetles. North American Beetle Fauna Project. The Biological Research Institute of America, Inc., July 1977, pp. 4.1-4.68.

Erwin, T.L., 1978. Studies of the subtribe Tachyina (Coleoptera: Carabidae: Bembidiini), Supplement C: Discovery of the sister group of Mioptachys Bates, in the Amazon Basin--Inpa psydroides a new genus and species. The Coleopterists Bulletin 32(1):29-36.

Erwin, T.L., 1978. The larva of Neotropical Enceladus gigas Bonelli (Coleoptera: Carabidae: Siagoninae: Enceladini) with notes on the phylogeny and classification of some of the more primitive tribes of ground beetles. The Coleopterists Bulletin 32(2):99-106.

Erwin, T.L., and G. N. House, 1978. A catalogue of the primary types of Carabidae (incl. Cicindelinae) in the collections of the United States National Museum of Natural History (USNM) (Coleoptera). The Coleopterists Bulletin 32(3):231-255.

Erwin, T.L., 1978. Systematic, natural history, and zoogeographic notes on the Genus Agra Fabricus, with a description of a new species from Panamá (Coleoptera: Carabidae: Lebiini). The Coleopterists Bulletin 32(4):261-268.

Erwin, T.L., and A. L. Halpern, 1978. Max Bänninger: His collection and publications (Coleoptera: Carabidae). The Coleopterists Bulletin 32(4):357-366.

Erwin, T.L., 1978. Techniques. The Coleopterists Bulletin 32(4):372.

Erwin, T.L., G. E. Ball, D. R. Whitehead, A. L. Halpern, (eds.), 1979. Carabid Beetles: Their evolution, natural history, and classification. Proceedings of the First International Symposium of Carabidology. Dr. W. Junk b.v. Publishers, The Hague. 635 pp.

Erwin, T.L., 1979. The American connection, past and present, as a model blending dispersal and vicariance in the study of biogeography, pp. 355-367. In, Erwin, T.L., G. E. Ball, D. R. Whitehead, A. L. Halpern, (eds.). Carabid Beetles: Their evolution, natural history, and classification. Proceedings of the First International Symposium of Carabidology. Dr. W. Junk b.v. Publishers, The Hague.

Erwin, T.L., 1979. A review of the natural history and evolution of ectoparasitoid relationships in carabid beetles, pp. 479-484. In, Erwin, T.L., G. E. Ball, D. R. Whitehead, A. L. Halpern, (eds.). Carabid Beetles: Their evolution, natural history, and classification. Proceedings of the First International Symposium of Carabidology. Dr. W. Junk b.v. Publishers, The Hague.

Erwin, T.L., 1979. Thoughts on the evolutionary history of ground beetles: Hypotheses generated from comparative faunal analyses of lowland forest sites in temperate and tropical regions, pp. 539-592. In, Erwin, T.L., G. E. Ball, D. R. Whitehead, A. L. Halpern, (eds.). Carabid Beetles: Their evolution, natural history, and classification. Proceedings of the First International Symposium of Carabidology. Dr. W. Junk b.v. Publishers, The Hague.

Erwin, T.L., 1980. Systematic and zoogeographic notes on Apotomini, an old world faunal element new to the western hemisphere list, with a description of a new species from Brazil (Coleoptera: Carabidae). The Coleopterists Bulletin 34(1):99-104.

Erwin, T.L., & D.H. Kavanaugh, 1980. On the identity of Bembidion puritanum Hayward (Coleoptera: Carabidae: Bembidiini). The Coleopterists Bulletin 34(2):241-242.

Erwin, T.L., and J.C. Scott, 1981. Seasonal and size patterns, trophic structure, and richness of Coleoptera in the tropical arboreal ecosystem: The fauna of the tree Luehea seemannii Triana and Planch in the Canal Zone of Panamá. The Coleopterists Bulletin 34(3):305-322.

Erwin, T.L., and D.H. Kavanaugh, 1981. Systematics and zoogeography of Bembidion Latreille: I. The carlhi and erasum groups of Western North America (Coleoptera: Carabidae: Bembidiini). Ent. Scand. Supplement 15:33-72.

Erwin, T.L., 1981. A synopsis of the immature stages of Pseudomorphini (Coleoptera: Carabidae) with notes on tribal affinities and behavior in relation to life with ants. The Coleopterists Bulletin 35(1):53-68.

Erwin, T.L., 1981. Taxon pulses, vicariance, and dispersal: An evolutionary synthesis illustrated by carabid beetles, pp. 159-183. In, Nelson, G. and D. Rosen, (eds.). Vicariance Biogeography: A Critique. Columbia University Press, New York.

Erwin, T.L., 1981. Natural history of Plummers Island, Maryland. XXVI. The ground beetles of a temperate forest site (Coleoptera: Carabidae): An analysis of fauna in relation to size, habitat selection, vagility, seasonality, and extinction. Bulletin of the Biological Society of Washington 5:105-224.

Erwin, T.L., 1981. Canopy beetles: Clarification of heat gain and water loss. The Coleopterists Bulletin 35(3):315-316.

Erwin, T.L., 1982. Agra, arboreal beetles of Neotropical forests: erythropus group systematics (Carabidae). Systematic Entomology 7:39-71.

Erwin, T.L., and J. Adis, 1982. Amazon inundation forests: Their role as short-term refuges and generators of species richness and taxon pulses, pp. 358-371. In, Prance, G. (ed.). Biological Diversification in the Tropics. Columbia University Press, New York.

Erwin, T.L., 1982. Agra, arboreal beetles of Neotropical forests: platyscelis group systematics (Carabidae). Systematic Entomology 7:185-210.

Erwin, T.L., 1982. Small terrestrial ground-beetles of Central America (Carabidae: Bembidiina and Anillina). Proceedings of the California Academy of Sciences 42(19):455-496.

Dritschilo, W. and Erwin, T.L., 1982. Responses in abundance and diversity of cornfield carabid communities to differences in farm practices. Ecology, 63(4):900-904.

Erwin, T.L., 1982. Tropical forests: Their richness in Coleoptera and other arthropod species. The Coleopterists Bulletin 36(1):74-75.

Erwin, T.L., 1983. Tropical forest canopies, the last biotic frontier. Bulletin of the Entomological Society of America 29(1):14-19.

Erwin, T.L., 1983. Baron Maximilien de Chaudoir: Chaudoir's studies of the taxa Agrina and Brachinitae (Coleoptera: Carabidae). The Coleopterists Bulletin 36(3):501-511.

Ball, G. E. and Erwin, T.L., 1983. The Baron Maximilien de Chaudoir: Inheritance, associates, travels, work, and legacy. The Coleopterists Bulletin 36(3):475-501.

Erwin, T.L., 1983. Agra, arboreal beetles of Neotropical forests: famula and formicaria groups systematics (Carabidae). Systematic Entomology 8:263-292.

Erwin, T.L., 1983. Beetles and other arthropods of the tropical forest canopies at Manaus, Brasil, sampled with insecticidal fogging techniques, pp. 59-75. In, Sutton, S. L., T. C. Whitmore, and A. C. Chadwick (eds.). Tropical Rain Forests: Ecology and Management. Blackwell Scientific Publications. Oxford, United Kingdom.

Erwin, T.L., 1984. Agra, arboreal beetles of Neotropical forests: palmata group systematics (Carabidae). Systematic Entomology 9:9-48.

Erwin, T.L., 1984. Studies of the Tribe Bembidiini (Coleoptera: Carabidae): Lectotype designations and species group assignments for Bembidion species described by Thomas L. Casey, et al. Pan-Pacific Entomologist 60(3):165-187.

Erwin, T.L., 1984. Composition and origin of the ground-beetle fauna of Sri Lanka (Coleoptera; Carabidae), pp. 371-389. In, Fernando, C. H. (ed.). Ecology and Biogeography of Sri Lanka. Dr. W. Junk b.v. Publishers, The Hague.

Erwin, T.L., and L.L. Sims, 1984. Carabid beetles of the West Indies (Insecta: Coleoptera): A synopsis of the genera and checklists of tribes of Caraboidea, and of the West Indian species. Quaestiones entomologicae 20:351-464.

Erwin, T.L., 1984. Small terrestrial ground beetles of the Amazon Basin (Coleoptera: Bembidiini: Tachyina and Anillina). Amazoniana 8(4):511-518.

Kavanaugh, D.H. and Erwin, T.L., 1985. Trechus obtusus Erichson (Coleoptera: Carabidae), a european ground beetle, on the pacific coast of North America: Its distribution, introduction and spread. Pan-Pacific Entomologist 61(2):170-179.

Lindroth, C. H., F. Bangsholt, R. Baranowski, T.L. Erwin, P. Jorum, B.-O. Landon, D. Refseth, and H. Silfverberg, 1985. The Carabidae (Coleoptera) of Fennoscandia and Denmark. Vol. 1. E. J. Brill Publishers, Copenhagen. 225 pp.

Erwin, T.L., 1985. Tambopata Reserved Zone, Madre de Dios, Perú: History and description of the reserve. Revista Peruana de Entomologia. 27:1-8.

Erwin, T.L., 1985. The taxon pulse: A general pattern of lineage radiation and extinction among carabid beetles, pp. 437-472. In, Ball, G. E. (ed.). Taxonomy, phylogeny, and zoogeography of beetles and ants: A volume dedicated to the memory of Philip Jackson Darlington Jr. 1904 - 1983. Dr. W. Junk b.v. Publishers, The Hague.

Erwin, T.L., and N. Stork, 1985. The Hiletini, an ancient and enigmatic tribe of Carabidae with a pantropical distribution (Coleoptera). Systematic Entomology 10:405-451.

Erwin, T.L., 1986. Agra, arboreal beetles of Neotropical forests: mixta-group, virgata-group, and ohausi-group systematics (Carabidae). Systematic Entomology 11:293-316.

Lindroth, C. H., F. Bangsholt, R. Baranowski, T.L. Erwin, P. Jorum, B.-O. Landin, D. Refseth, and H. Silfverberg, 1986. The Carabidae (Coleoptera) of Fennoscandia and Denmark. Vol. 2. E. J. Brill Publishers, Copenhagen. 497 pp.

Adis, J., W. Paarmann and T.L. Erwin, 1986. On the natural history and ecology of small terrestrial Ground-Beetles (Col.: Bembidiini: Tachyina: Polyderis) from an amazonian black-water inundation forest (Satz) pp. 413-427. In, den Boer, et al. (eds.). Carabid Beetles. Gustav Fischer, Stuttgart, New York.

Erwin, T.L., 1987. Agra, arboreal beetles of Neotropical forests: feisthameli group systematics (Carabidae). Systematic Entomology 12:137-161.

Erwin, T.L., 1988. The tropical forest canopy: the heart of biotic diversity, pp. 123-129. In, Wilson, E. O. (ed.). Biodiversity. National Academy Press, Washington, D.C.

Erwin, T.L., and M.G. Pogue, 1988. Agra, arboreal beetles of Neotropical forests: Biogeography and the forest refugium hypothesis (Carabidae), pp. 161-188. In, Heyer, W. R. and P. E. Vanzolini (eds.). Neotropical Distribution Patterns: Proceedings of a 1987 Workshop. Proceedings of the Brasilian Academy of Sciences.

Farrell, B.D. and T.L. Erwin, 1988. Leaf-beetles (Chrysomelidae) of a forest canopy in amazonian Perú: synoptic list of taxa, seasonality and host-affiliations, pp. 73-90. In, Jolivet, P., E. Petitpierre, and T. Hsiao, (eds.). The Biology of the Chrysomelidae. Dr. W. Junk b.v. Publishers, The Hague.

Erwin, T.L., 1989. Canopy arthropod biodiversity: A chronology of sampling techniques and results. Revista Peruana de Entomologia 32:71-77.

Erwin, T.L., 1991. The ground-beetles of Central America (Carabidae) I: Carabinae (in part): Notiophilini, Loricerini, Carabini. Smithsonian Contributions to Zoology No. 501, 30 pp.

Kavanaugh, D.H. and T.L. Erwin, 1991. The tribe Cicindini Bänninger (Coleoptera: Carabidae): Comparative morphology, natural history, and reclassification. Proceeding of the Entomological Society of Washington 93(2):356-389.

Erwin, T.L., 1991. An evolutionary basis for conservation strategies. Science 253:750-752.

Erwin, T.L., 1991. How many species are there?: Revisited. Conservation Biology 5(3):330-333.

Erwin, T.L., 1991. Establishing a tropical species co-occurrence database. Part 1. A plan for developing consistent biotic inventories in temperate and tropical habitats. Memoria del museo de historia natural, No. 20, pp. 1-16.

Erwin, T.L., and M. Kabel, 1991. Establishing a tropical species co-occurrence database. Part 2. An automated system for mapping dominant vegetation. Memorias del museo de historia natural, No. 20, pp. 17-36.

Erwin, T.L., and D.L. Pearson, 1991. Establishing a tropical species co-occurrence database. Part 3. An integrated approach toward understanding biological diversity. Memorias del museo de historia natural, No 20, pp. 37-45.

Erwin, T.L., 1991. The significance of diversity: New challenges for the entomologist, pp. 25-39. In, Vinson, S. B. and R. L. Metcalf, (eds.). Entomology serving society: Emerging technologies and challenges. Entomological Society of America, Lanham, Maryland.

Erwin, T.L., 1991. Natural history of the carabid beetles at the BIOLAT Rio Manu Biological Station, Pakitza, Perú. Revista Peruana de Entomologia, 33:1-85.

Erwin, T.L., 1992. A current vision of insect diversity, pp. 91-97. In, Sarukhán, J., and R. Dirzo, (eds.). México ante los retos de la biodiversidad, Mexico confronts the challenges of biodiversity. Comisión Nacional para el Conociniento y Uso de la Biodiversidad, Mexico City.

Erwin, T.L., 1992. Life in the Treetops, Australian Encyclopedia of Life (1 page). Australia.

Kavanaugh, D. H. and T.L. Erwin, 1992. Extinct or Extant? A new species of intertidal bembidiine (Coleoptera: Carabidae: Bembidiini) from the Palos Verdes Peninsula, California. The Coleopterists Bulletin 46(3): 311-320.

Kremen, C., R. K. Colwell, T.L. Erwin, D. D. Murphy, R. F. Noss and M. A. Sanjayan, 1993. Terrestrial arthropod assemblages: Their use in conservation planning. Conservation Biology 7(4):796-808.

Erwin, T.L., 1993. Stratification and distribution of neotropical forest microhabitats and associated arthropod faunules; a methodology report. Selbyana 14:8.

Erwin, T.L., 1994. Arboreal beetles of tropical forests: The Xystosomi group, subtribe Xystosomina (Coleoptera: Carabidae: Bembidiini). Part I. Character analysis, taxonomy, and distribution. The Canadian Entomologist 126(3):549-666. 155 figs, 2 tables.

Erwin, T.L., 1995. Measuring arthropod biodiversity in the tropical forest canopy, pp 109-127. In, M. Lowman and N. Nadkarni (eds.). Forest Canopies, Academic Press Inc.

Erwin, T.L., 1996. Arboreal Beetles of Neotropical Forests: Agra Fabricius, the cayennensis complex (Coleoptera: Carabidae: Lebiini: Calleidina). Annales Zoologici Fennici 33(1):17-21. 2 figs.

Erwin, T.L., 1993. Agra, arboreal beetles of Neotropical forests: rufoaenea and quararibea group systematics (Carabidae). Revista peruana de Entomologia, (1991) 34:15-28. 34 figs.

Erwin, T.L., 1996. Biodiversity at its utmost: Tropical forest beetles, pp. 27-40. In, Reaka-Kudla, M. L., D. E. Wilson and E. O. Wilson (eds), Biodiversity II. Understanding and protecting our biological resources. Joseph Henry Press, Washington, D. C.

Venable, G.L. & T.L. Erwin, 1997. Mapping of the biodiversity site at Pakitza, pp. 29-42. In, Wilson, D. E. and A. Sandoval (eds.), Manu, The biodiversity of southeastern Peru. Editorial horizonte, Lima.

Erwin, T.L., 1997. Natural history of the carabid beetles at the BIOLAT Biological Station, Rio Manu, Pakitza, Perú, Supplement I. Additional records, pp. 359-368. In, Wilson, D. E. and A. Sandoval (eds.), Manu, The biodiversity of southeastern Peru. Editorial horizonte, Lima.

List of per-reviewed articles reprinted in books, magazines, journals:

1993. [Erwin, T.L., 1991. An evolutionary basis for conservation strategies. Science 253:750-752] In, Perspectives on Biodiversity: Case studies of genetic resource conservation and development (eds. C.S. Potter, J.I. Cohen, D. Janczewski) AAAS, pp 19-25.

SINCE LAST REVIEW PERIOD

Adis, J, M.A. Amorim, T.L. Erwin, & T. Bauer, 1997 (1998). On Ecology, life history and survival strategies of a wing-dimorphic ground beetle (Col. Carabidae: Odacanthini: Colliuris) inhabiting central Amazonian inundation forests. Studies on Neotropical Fauna and Environment. 32:174-192 [provided significant input - identification and descriptions of the new species]

Kress, J. ¼..T.L. Erwin, et al. 1998. Amazon biodiversity: assessing conservation priorities with taxonomic data. Biodiversity and Conservation 7:1577-1587. [provided data, reviewed ms]

Erwin, T.L., & D.H. Kavanaugh, 1999. Studies of the Subtribe Tachyina (Coleoptera: Carabidae: Bembidiini), Supplement D: Description of a Male of Costitachys inusitatus Erwin, with Notes on Distribution of this Species. The Coleopterists Bulletin 53(1):52-55. [wrote the paper]

Erwin, T.L., 1998. Evolution at the equator arboreal and alticolous beetles and their taxon pulses with descriptions of a new Agra subclade and its species (Coleoptera: Carabidae: Lebiini) pp 491-510. In, Phylogeny and classification of Caraboidea (Coleoptera: Adephaga). Proceedings of a Symposium (28 August, 1996, Florence, Italy), XX International Congress of Entomology (G.E. Ball, A. Casale & A. Vigna Taglianti, Editors). Museo Regionale Scienze Naturali- Torino, Atti. Torino.

Erwin, T.L., 2000. Arboreal beetles of Neotropical forests: Agra Fabricius, the Novaurora (Coleoptera: Carabidae: Lebiini: Calleidina). Smithsonian Contributions to Zoology, No. 608, pp. i-iv, 1-33.

Erwin, T.L., & D.H. Kavanaugh, 2000. A New Species of the Tachyta Subgenus Paratachyta Erwin (Coleoptera: Carabidae: Bembidiini: Tachyina) from Southeastern China with a New Key and Analysis of Phylogenetic Relationships among Species of the Subgenus, Pp. 183-194. In, Zamotajlov, A. & R. Sciaky (eds.). Advances in carabidology (Papers dedicated to the memory of Prof. Oleg L. Kryzhanovskij). MUISO publications, Krasnodar, 473pp. [wrote the penultimate draft]

Erwin, T.L., 2000. Arboreal beetles of Neotropical Forests: Taxonomic supplement for the platyscelis group with new species and distribution records (Coleoptera:Carabidae, Lebiini, Agrina). The Coleopterists Bulletin 54(1):90-119.

Heyer, W.R., ¼. T.L. Erwin, et al. 2000. Amazonian biotic data and conservation decisions. Ciência e Cultura 51(5,6):372-385. [provided data, reviewed ms]

Erwin, T.L., 2000. Arboreal beetles of Neotropical Forests: Taxonomic supplement for the Agra virgata and ohausi groups with a new species and additional distribution records (Coleoptera:Carabidae). The Coleopterists Bulletin 54(2):251-262

Erwin, T.L. & P.J. Johnson, 2000. Naming species, a new paradigm for crisis management in taxonomy: Rapid journal validation of scientific names enhanced with more complete descriptions on the internet. The Coleopterists Bulletin 54(3):269-278. [wrote the paper]

Erwin, T.L., 2000. A new genus and species of Lachnophorini and two new species of Lebiini from Costa Rica (Coleoptera: Carabidae). The Coleopterists Bulletin 54(3):279-283.

Erwin, T.L., 2000. Lumpers and Splitters. Whole Earth Magazine. Fall, 2000, pp. 45-46.

Erwin, T.L., 2000. Typecast: The Codes of Nomenclature. Whole Earth Magazine. Fall, 2000, p. 47.

Erwin, T.L., 2001. Forest Canopies, Animal Diversity. Encylopedia of Biodiversity 3:19-25. Academic Press.

Arndt, E., S. Kimse, & T.L. Erwin, 2001. Arboreal beetles of Neotropical forests: Agra Fabricius, larval descriptions with notes on natural history and behaviour (Coleoptera, Carabidae, Lebiini, Agrina). The Coleopterists Bulletin 55(3):297-311. [rewrote the paper, described the larvae, illustrations, etc.]

Lucky, A., T.L. Erwin, and J.D. Witman, 2002. Temporal and Spatial Diversity and Distribution of Arboreal Carabidae (Coleoptera) in a Western Amazonian Rain Forest. Biotropica 34(3):376–386. [provided all data, field work, helped this student with her Senior Thesis analysis]

Erwin, T.L. 2002. The Beetle Family Carabidae of Costa Rica: Twenty-nine new species of Costa Rican Agra Fabricius 1801 (Coleoptera:Carabidae, Lebiini, Agrina). Zootaxa 119:1-68.

Philips, O. … T.L. Erwin, et al. 2002. Increasing dominance of large lianas in Amazonian forests. Nature 418, 770-774. [provided data, reviewed the ms]

Alvarez, M., K. Balta, L. Campos, C. del Aguila, T. P. Feria, L. Nolivos & Erwin, T.L. (2002) ¿Qué hay mas allá del orificio?: riqueza y abundancia de los insectos acuáticos presentes en los internodos del bambú Guadua weberbaueri. In, G. Servat & D. Cadena, eds., Ecología de Ecosystemas Amazónicos, 2002-13, 88-91. [structured the research, reviewed the paper]

Feria, A. T. P., Balta, K. & Erwin, T.L. (2002). Efecto del volúmen de agua sobre la riqueza de invertebrados presente en los internodos del bambu Guadua weberbaueri. In, G. Servat & D. Cadena, eds., Ecología de Ecosystemas Amazónicos, 2002-13, 100-102. [structured the research, reviewed the paper]

Aschero, V., Zenteno, L., Rios, S., Gonzáles, C., Achicahuala, J., Vargas, J. & Erwin, T.L. (2003) La misma obra, diferentes actores: Estructura y composición de las comunidades de insectos acuáticos en los [sic] internudos del bambú (Guadua weberbaueri). In, G. Servat, M. Rodriguez, & D. Cadena, eds., Ecología de Ecosystemas Amazónicos, 2003-13, 87-90. [structured the research, reviewed the paper]

Delgado, C., Costantino, M., Curo, A., Estrada, N., Fagua, C., Machuca, J., & Erwin T.L. (2003) Especie clave para el establecimiento de fauna acuática en los [sic] internados de bambú: ¿Escarabajo o Carpintero? In, G. Servat, M. Rodriguez, & D. Cadena, eds., Ecología de Ecosystemas Amazónicos, 100-102. [structured the research, reviewed the paper]

Erwin, T.L. & R.F. Medina, 2003. The Neotropical species Askalaphium depressum (Bates): Larval description, first diagnosis and illustrations of immature Ctenodactylini, with natural history notes on the genus and tribe (Coleoptera: Carabidae). Zootaxa 273:1-15. [wrote the paper]

Erwin, T.L., 2004. The Beetle Family Carabidae of Costa Rica and Panamá: Descriptions of four new genera and six new species with notes on their way of life (Insecta: Coleoptera). Zootaxa 537:1-18.

Erwin, T.L. & V. Aschero, 2004. Cicindis horni Bruch (Coleoptera: Carabidae, Cicindini): The Fairy Shrimp Hunting Beetle, its way of life on the Salinas Grandes of Argentina. Zootaxa 553: 1-16. [wrote the paper]

Erwin, T.L. (2004) The Biodiversity Question: How many species of terrestrial arthropods are there? Pp. 259-269. In, Lowman, M. & Brinker (eds.) Forest Canopies, 2nd Edition, Academic Press.

Erwin, T.L. (2004) The Beetle Family Carabidae of Costa Rica: The Genera of the Cryptobatida Group of Subtribe Agrina, Tribe Lebiini, with new species and notes on their way of life (Insecta: Coleoptera). Zootaxa, 662: 1-54.

Erwin, T.L., 2004.The Beetle Family Carabidae of Costa Rica: The Genus Epikastea Liebke of the Plochonida Group, with new Neotropical species and notes on their way of life (Insecta: Coleoptera, Lebiini, Agrina). Zootaxa, 790: 1-20.

Erwin, T.L., M.C. Pimienta, O.E. Murillo & V. Aschero 2004. Mapping patterns of β-diversity for beetles across the western Amazon Basin: A preliminary case for improving conservation strategies. Proceedings of the California Academy of Sciences, ser. 4, 56(Suppl. I) No. 7; 72–85. [wrote the paper]

Philips, O.L. … Erwin, T.L. (2005) Late twentieth-century patterns and trends in Amazon tree turnover. Pp. 107-128. In, Yadvinders, M. & Oliver, O.L. (Eds.) Tropical Forests & Global Atmospheric Change, 260pp. Oxford University Press, UK. [provided data, reviewed the ms]

Baker, T.R., Philips, O.L. … Erwin, T.L. (2005) Late twentieth-century trends in the biomass of Amazonian forest plots. Pp. 129-141. In, Yadvinders, M. & Oliver, O.L. (Eds.) Tropical Forests & Global Atmospheric Change, 260pp. Oxford University Press, UK. [provided data, reviewed the ms]

Aschero, V., Zenteno, L., Rios, S., Gonzáles, C., Achicahuala, J., Vargas, J. & Erwin, T.L. (2003) La misma obra, diferentes actores: Estructura y composición de las comunidades de insectos acuáticos en los [sic] internudos del bambú (Guadua weberbaueri). Ecología de Ecosystemas Amazónicos, 2003-13, 87-90.

Delgado, C., Costantino, M., Curo, A., Estrada, N., Fagua, C., Machuca, J., & Erwin T.L. (2003) Especie clave para el establecimiento de fauna acuática en los [sic] internados de bambú: ¿Escarabajo o carpintero? Ecología de Ecosystemas Amazónicos, 2003-13, 100-102.

Feria, A. T. P., Balta, K. & Erwin, T.L. (2002). Efecto del volúmen de agua sobre la riqueza de invertebrados presente en los internodos del bambu Guadua weberbaueri. Ecología de Ecosystemas Amazónicos, OTS, 2002-13, 100-102.

Alvarez, M., K. Balta, L. Campos, C. del Aguila, T. P. Feria, L. Nolivos & Terry L. Erwin. 2002. ¿Qué hay mas allá del orificio?: riqueza y abundancia de los insectos acuáticos presentes en los internodos del bambú Guadua weberbaueri. Ecología de Ecosystemas Amazónicos, OTS, 2002-13, 100-102.

Erwin, T.L. & Moore, W. (2007) The Beetle Family Carabidae of Costa Rica: The Genus Moriosomus Motschulsky with a new Neotropical species and notes on their way of life (Insecta: Coleoptera, Carabidae, Moronini). Zootaxa, 1438: 49-63.

Steiner, W.E. Jr.,  Erwin, T.L. & Ward, R. (2007) Phloeoxena signata (Dejean): A northern range extension to Maryland and Tennessee, USA, and the first record for Costa Rica (Coleoptera: Carabidae). The Coleopterists Bulletin, 61(2):224-226.

List of per-reviewed articles reprinted in books, magazines, journals:

Erwin, T.L., 1982. Tropical forests: Their richness in Coleoptera and other arthropod species. The Coleopterists Bulletin 36(1):74-75. In, 2002. Chazdon, R.L. & T.C. Whitmore (eds.) Foundations of Tropical Forest Biology: Classic Papers with Commentaries. The University of Chicago Press. 862 pp.

List of books:

Lindroth, C. H., F. Bangsholt, R. Baranowski, T.L. Erwin, P. Jorum, B.-O. Landon, D. Refseth, and H. Silfverberg, 1985. The Carabidae (Coleoptera) of Fennoscandia and Denmark. Vol. 1. E. J. Brill Publishers, Copenhagen. 225 pp.

Lindroth, C. H., F. Bangsholt, R. Baranowski, T.L. Erwin, P. Jorum, B.-O. Landin, D. Refseth, and H. Silfverberg, 1986. The Carabidae (Coleoptera) of Fennoscandia and Denmark. Vol. 2. E. J. Brill Publishers, Copenhagen. 497 pp.

Adis, J. (Scientific Editor), T.L. Erwin (Scientific Coordinator) 1988. Translation: Lindroth, C. H., The Ground Beetles (Carabidae) of Fennoscandia: A zoogeographic study. Part II. Maps. Amerind Publishing Co. Pvt. Ltd., New Delhi, India. 271

Adis, J. (Scientific Editor), T.L. Erwin (Scientific Coordinator) 1993. Translation: Lindroth, C. H., The Ground Beetles (Carabidae) of Fennoscandia: A zoogeographic study. Part I. Specific Zoogeographic Facts. Amerind Publishing Co. Pvt. Ltd., New Delhi, India. 630 p.

Adis, J. (Scientific Editor), T.L. Erwin (Scientific Coordinator) 1993. Translation: Lindroth, C. H., The Ground Beetles (Carabidae) of Fennoscandia: A zoogeographic study. Part III. General Facts. Amerind Publishing Co. Pvt. Ltd., New Delhi, India. 814 p.

Erwin, T.L., (ed.). 1991. Establishing a tropical species co-occurrence database. Memoria del museo de historia natural, No. 20, pp. 1-45.

List of Reviews

Erwin, T.L., 1972. The Carabid Beetles of New Guinea, by Philip J. Darlington. Systematic Zoology 21(3):343-347.

Erwin, T.L., 1974. Surtsey, Iceland: The Development of a New Fauna, 1963-1970, Terrestrial Invertebrates, by Carl H. Lindroth, Hugo Andersson, Hogni Bodvarsson, and Sigurdur H. Richter. Entomologica Scandinavica, Suppl. 5, 1973. Munksgaard, International Booksellers and Publishers Ltd., 35, Norre Sogade, DK-1370 Copenhagen K, Denmark. 280 p.; 50 fig. Paper 40 Danish Crowns ($5.35). The Coleopterists Bulletin 28(3):166.

Erwin, T.L., 1978. Handbook of Common New Guinea Beetles, by J. Linsley Gressitt and Richard W. Hornabrook, 1977, Wau Ecology Institute, Box 77, Wau, Papua New Guinea, viii + 87 pp., with 35 figs. US $4.50, AU $3.50, K 3.00. ESA Bulletin 24(4):443-444.

Erwin, T.L., 1991. Four Neotropical Rainforests, by A. H. Gentry, (ed.), 1990. Yale University Press, New Haven and London, i-xiii, 1-627 pp. Association of Systematic Collections Newsletter 19(3):38-40.

Erwin, T.L., 1993. Conserving Biodiversity: A Research Agenda for Development Agencies. Report of a Panel of the Board on Science and Technology for International Development, U. S. National Research Council. National Academy Press, 1992. 116 pp. In, The Quarterly Review of Biology 66(2):300.

Erwin, T.L., 1994. Insects of Panamá and Mesoamerica: Selected Studies, by D. Quintero and A. Aiello, 1992. Oxford Science Publications. 692 pp, The Quarterly Review of Biology 69(3):410.

Erwin, T.L., 1998. Forests and Insects. Allan D. Watt, Nigel E. Stork, and Mark D. Hunter (editors) London: Chapman & Hall, 1997 pp. i-xv, 1-406. Biodiversity and Conservation 7(12):1662-1664.

Erwin, T.L., 1997. Timothy Swanson, Global Action for Biodiversity, An International Framework for implementing the convention on biological diversity. Earthscan Publications Ltd. London, 191 pp. ISBN: 1 85383 358 4.

Erwin, T.L. 2003. Review: The Insects of Hawaii, Volume 16, Hawaiian Carabidae (Coleoptera), Part 1: Introduction and Tribe Platynini, J.K. Liebherr and E.C. Zimmerman, University of Hawai’i Press, Honolulu, 2000, 494 pages, ISBN 0-8248-2356-7, In, Annals of the Entomological Society of America 96(4):596.

List of popular articles:

Erwin, T.L., 1976. Chirp and rattle, dart and glitter! Smithsonian magazine, November 1976, pp. 116-120, (article on the Insect Zoo, photographs by Kjell Sandved).

Erwin, T.L., 1995. Sierrazul, Ecuador: A primeval cloud forest overlooking the amazon basin (ready for biotic researchers and conservation biologists). Newsletter of the International Center of Tropical Ecology 2(2):6-7.

Erwin, T.L. (2005) Roy A. Crowson. Founders’ Memorial Award: the Entomologist we honor. The 53rd Annual Meeting of the Entomological Society of America, Ft. Lauderdale, Program Booklet, pp. 15-16.

List of electronic media:

Erwin, T.L., 1996. Checklist of the Western Hemisphere Caraboidea (COLEOPTERA), 12Mb. [Includes all 12,000 species’ distributions and taxonomy]

On, NMNH Gopher; http://nmnhwww.si.edu/gopher-menus/

[Updated, 2001] Checklist of the Western Hemisphere Caraboidea (COLEOPTERA).
http://entomology.si.edu/Entomology/WstrnCarabids/search.lasso

Erwin, T.L., 2000. Illustrated guide to Carabidae genera (COLEOPTERA).
http://entomology.si.edu/Entomology/CarabidGenus/genweb.lasso

Erwin, T.L. & R. Medina, 2000. Plummers Island Natural History
http://entomology.si.edu/Entomology/plummers/plummers.lasso

Moore, W., T.L. Erwin, and D.H. Kavanaugh 2002. The Family Carabidae: Introduction. http://www.inbio.ac.cr/papers/carabidae/esp/index.html

Erwin, T.L., D.H. Kavanaugh, and W. Moore 2002. The Family Carabidae: Classification of the Costa Rican Taxa. http://www.inbio.ac.cr/papers/carabidae/esp/clasificacion.htm

Erwin, T.L., D.H. Kavanaugh, and W. Moore 2002. The Family Carabidae: Key to the Tribes and Genera known to occur in Costa Rica. http://www.inbio.ac.cr/papers/carabidae/esp/images/CR_carabid_key.doc

Revised, July 2004.

Erwin, T.L., 2000. A new genus and species of Lachnophorini from Costa Rica (Coleoptera: Carabidae, Quammenis spectablis). http://www.inbio.ac.cr/

Erwin, T.L., 2000. A new species of Chelonodema from Costa Rica (Coleoptera: Carabidae, Lebiini). http://www.inbio.ac.cr/

Erwin, T.L., 2000. A new species of Agra from Costa Rica (Coleoptera: Carabidae, Lebiini). http://www.inbio.ac.cr

e-Abstract: Erwin, T.L., 2003. Mapping patterns of β-diversity for beetles across the western Amazon Basin: A preliminary case for improving conservation strategies. 4th Annual Ecological Integration Symposium "Natural History and Modern Conservation" Texas A & M University.
http://eco-symposium.tamu.edu/wfscnet/groups/symposium/erwin.htm

Abstracts:

Erwin T.L. 1999. Mosquitoes, birds, bats, beetles & other flying and nonflying things: How instructive are their distribution for tropical conservation strategies? AMCA Annual Meeting Symposium, “Species diversity & challenges in systematics” St. Louis, MO

Erwin T.L. 2000. Canopy Arthropods of the Western Amazon Basin. XXI ICE Symposium: Amazon Insects. Abstract Book I: 98.

Erwin T.L. 2001. A Taxonomist’s Vision: Knowing all Earth’s species (Biodiversity and complementarity underpinning conservation strategies). Proceedings of the Annual Meeting of the Bahamian Natural History Field Station, San Salvador, Bahamas.

Erwin T.L. 2003. Mapping patterns of β-diversity for beetles across the western Amazon Basin: A preliminary case for improving conservation strategies. 84th Annual Meeting of the Pacific Division, AAAS. San Francisco, CA.

Erwin T.L. 2005. Artrópodos del dosel en el oeste de la Cuenca del Amazonas. In, Libro de resúmenes, Congreso Argentino de Entomología, 2005, San Miguel de Tucumán, Argentina.

Erwin T.L. 2006. Monitoring changes in canopy arthropod populations through time in the western Amazon Basin, Yasuni area, Ecuador. Joint Meeting of the Entomological Society of Canada and the Société d’entomologie du Quebéc, Montreal, Canada, November 2006, Ecology Symposium.

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